[1]

Gáperová S, Gáper J, Gašparcová T, Náplavová K, Pristaš P. 2016. Morphological variability of Fomes fomentarius basidiomata based on literature data. Annales Universitatis Paedagogicae Cracoviensis Studia Naturae 1(1):42−51

[2]

Mukhin VA, Zhuykova EV, Badalyan SM. 2018. Genetic variability of the medicinal tinder bracket polypore, Fomes fomentarius (Agaricomycetes), from the Asian part of Russia. International Journal of Medicinal Mushrooms 20(6):561−568

doi: 10.1615/intjmedmushrooms.2018026278
[3]

Peintner U, Kuhnert-Finkernagel R, Wille V, Biasioli F, Shiryaev A, et al. 2019. How to resolve cryptic species of polypores: an example in Fomes. IMA Fungus 10:17

doi: 10.1186/s43008-019-0016-4
[4]

Badalyan SM, Zhuykova EV, Mukhin VA. 2022. The phylogenetic analysis of Armenian collections of medicinal tinder polypore Fomes fomentarius (Agaricomycetes, Polyporaceae). Italian Journal of Mycology 51:23−33

doi: 10.6092/issn.2531-7342/14474
[5]

Badalyan SM, Zhuykova EV, Mukhin VA. 2026. The eco-geographical and phylogenetic analysis of Fomes fomentarius sensu lato (Polyporales, Agaricomycetes) in Armenia. Italian Journal of Mycology 55:8−23

doi: 10.60923/issn.2531-7342/22982
[6]

Buchalo AS. 1988. Higher Edible Basidiomycetes in Pure Culture. Kiev, Ukraine: Naukova Dumka

[7]

Buchalo AS, Wasser SP, Mikchaylova OB. 2011. Micromorphological characteristics of edible and medicinal macromycetes in pure culture. In Biological Characteristics of Medicinal Macromycetes in Culture, Vol 1. eds. Buchalo AS, Babitskaya WG, Bisko NA, Wasser SP, Dudka IA, et al. Kiev, Ukraine: Alterpress. pp. 105–134 (in Russian)

[8]

Badalyan SM, Sakeyan CZ. 2004. Morphological, physiological, and growth characteristics of mycelia of several wood-decaying medicinal mushrooms (Aphyllophoromycetideae). International Journal of Medicinal Mushrooms 6:347−360

doi: 10.1615/IntJMedMushr.v6.i4.50
[9]

Badalyan SM, Hughes KW, Sakeyan CZ, Helmbrecht E. 2006. Morphology, growth characteristics, and genetic variability of the edible medicinal mushroom Flammulina velutipes (W. Curt.: Fr.) singer collections. International Journal of Medicinal Mushrooms 8(3):263−278

doi: 10.1615/IntJMedMushr.v8.i3.80
[10]

Badalyan SM, Navarro-González M, Kües U. 2011. Taxonomic significance of anamorphic characteristics in the life cycle of coprinoid mushrooms. Proceedings of the 7th International Conference on Mushroom Biology and Mushroom Products 7 (ICMBMP7). 4−7 October, 2011, Arcachon, France. Vol. 1. Bordeaux: INRA. pp. 140−154 www.researchgate.net/publication/229163985

[11]

Badalyan SM, Szafranski K, Hoegger PJ, Navarro-González M, Majcherczyk A, et al. 2011. New Armenian wood-associated coprinoid mushrooms: Coprinopsis strossmayeri and Coprinellus aff. radians. Diversity 3:136−154

doi: 10.3390/d3010136
[12]

Badalyan SM, Gharibyan NG, Iotti M, Zambonelli A. 2012. Morphological and genetic characteristics of different collections of Ganoderma P. Karst. species. Proceedings of the 18th Congress of International Society for Mushroom Science, eds. Zhang J, Wang H, Chen M. Beijing: China Agriculture Press. pp. 247–254

[13]

Badalyan SM, Shnyreva AV, Iotti M, Zambonelli A. 2015. Genetic resources and mycelial characteristics of several medicinal polypore mushrooms (Polyporales, Basidiomycetes). International Journal of Medicinal Mushrooms 17:371−384

doi: 10.1615/intjmedmushrooms.v17.i4.60
[14]

Badalyan SM, Gharibyan NG, Iotti M, Zambonelli A. 2019. Morphological and ecological screenings of different collections of medicinal white-rot bracket fungus Ganoderma adspersum (Schulzer) Donk (Agaricomycetes, Polyporales). Italian Journal of Mycology 48:1−15

doi: 10.6092/issn.2531-7342/9092
[15]

Badalyan SM, Shahbazyan TA, Gharibyan NG. 2019. The morphological observation of mycelia of several Armenian strains of medicinal bracket fungus Fomes fomentarius (L.) Fr. (Polyporales, Agaricomycetes). Proceedings of the YSU B: Chemical and Biological Sciences 53(2):91−96

doi: 10.46991/PYSUB.2019.53.2.091
[16]

Badalyan SM, Borhani A. 2019. Morphological and growth characteristics of mycelial collections of medicinal xylotrophic mushrooms (Agaricomycetes) distributed in northern forests of Iran. Proceedings of the YSU B: Chemical and Biological Sciences 53(2):97−106

doi: 10.46991/PYSUB.2019.53.2.097
[17]

Kües U, Badalyan SM, Gießler A, Dörnte B. 2016. Asexual sporulation in agaricomycetes. In The Mycota: Growth, Differentiation, and Sexuality, Vol. I, 3rd Edition, ed. Wendland J. Cham: Springer. pp. 269–328 doi: 10.1007/978-3-319-25844-7_12

[18]

Kües U, Subba S, Kapli H, Khonsuntia W, Badalyan SM, et al. 2026. Ecology and genetics in asexual sporulation in agaricomycetes. In The Mycota: Growth, Differentiation and Sexuality, vol 1. ed. Wendland J. Cham: Springer. pp 241–336 doi: 10.1007/978-3-032-15548-1_11

[19]

Dresch P, D'Aguanno MN, Rosam K, Grienke U, Rollinger J, et al. 2015. Fungal strain matters: colony growth and bioactivity of the European medicinal polypores Fomes fomentarius, Fomitopsis pinicola, and Piptoporus betulinus. AMB Express 5:4

doi: 10.1186/s13568-014-0093-0
[20]

Zhuykova EV, Mukhin VA. 2022. Diversity and ecological features of phylogenetic lineages of tinder fungus in the Urals. Russian Journal of Ecology 53:366−372

doi: 10.1134/S1067413622050113
[21]

Gáperová S, Gáper J, Gallay I, Pristaš P, Slobodník B. 2025. Spatial distribution and host preferences of Fomes fomentarius and F. inzengae in Europe: a review. Folia Oecologica 52(2):202−218

doi: 10.2478/foecol-2025-0019
[22]

Garrido-Benavent I, Velasco-Santos JM, Pérez-De-Gregorio MȦ, Pasaban PM. 2020. Fomes inzengae (Ces. & De Not.). Cooke en la Península Ibérica. Butlletí Societat Micològica Valenciana 24:151−170 (in Spanish)

[23]

Tomšovský M, Kaeochulsri S, Kudláček T, Dálya LB. 2023. Ecological, morphological and phylogenetic survey of Fomes fomentarius and F. inzengae (Agaricomycetes, Polyporaceae) co-occurring in the same geographic area in Central Europe. Mycological Progress 22:79

doi: 10.1007/s11557-023-01928-y
[24]

Bondarzew AS. 1953. The Polyporaceae of the European USSR and Caucasus. Moscow: Academy of Sciences of the USSR (in Russian)

[25]

Gilbertson RL, Ryvarden L. 1987. North American Polypores. Abortiporus-Lindtneria. Vol. 1: Abortiporus − Lindtneria. Oslo: Fungiflora A/S. doi: 10.1002/jobm.3620270513

[26]

Cui YJ, Liu HG, Dai YC, Wang CG. 2024. A new species of Fomes (Polyporaceae, Basidiomycota) from southwestern China. Phytotaxa 661(3):282−292

doi: 10.11646/phytotaxa.661.3.5
[27]

Melik-Khachatryan JH, Martorosyan SN. 1971. Mycoflora of Armenian SSR. Vol. 2. Gasteromycetes and Aphyllophorales. Yerevan: YSU Press. (in Russian) http://publishing.ysu.am/hy/1458647238

[28]

Nobles MK. 1965. Identification of culture of wood-inhabiting Hymenomycetes. Canadian Journal of Botany 43:1097−1139

doi: 10.1139/b65-126
[29]

Stalpers JA. 1978. Identification of wood-inhabiting Aphyllophorales in pure culture. Studies in Mycology 16:1–248

[30]

Tiberius B, Cătălin T. 2012. Culture description of some spontaneous lignicolous macromycetes species. Journal of Plant Development 19:83−97

[31]

Badalyan SM, Gharibyan NG. 2024. Morpho-ecological peculiarities of mycelia of several hymenochaetoid medicinal mushrooms. In Current Mycology in Russia: Proceedings of the International Mycological Forum. eds. Sergeev YV, Kurakov AV. Moscow, Russia: National Academy of Mycolog (in Russian)

[32]

Gharibyan NG, Gasparyan AK, Badalyan SM. 2024. Study of antifungal activity of mycelia of Armenian collections of polypore mushroom Fomes inzengae (Agaricomycetes) toward dermatophytes. In Advances in Medical Mycology. Vol. 26, ed. Sergeev YV. Moscow, Russia: National Academy of Mycology. pp. 20–22 (in Russian)

[33]

Gharibyan NG, Barkhudaryan AL, Badalyan SM. 2025. Study of antibacterial activity of mycelia of bracket fungi Fomes fomentarius s. s. and F. inzengae (Polyporales, Agaricomycetes). In Advances in Medical Mycology, ed. Sergeev YV. Vol. 27. Russia, Moscow: National Academy of Mycology. pp. 318–322 (in Russian)

[34]

Badalyan S. 2012. Medicinal aspects of edible ectomycorrhizal mushrooms. In Edible Ectomycorrhizal Mushrooms, eds. Zambonelli A, Bonito GM. Heidelberg, Berlin, Heidelberg: Springer. pp. 317–334 doi: 10.1007/978-3-642-33823-6_18

[35]

Badalyan SM, Barkhudaryan A, Rapior S. 2019. Recent progress in research on the pharmacological potential of mushrooms and prospects for their clinical application. In Medicinal Mushrooms − Recent Progress in Research and Development, eds. Agrawal DC, Dhanasekeran M. Singapore: Springer Nature. pp. 1–70 doi: 10.1007/978-981-13-6382-5_1

[36]

Badalyan SM, Morel S, Barkhudaryan A, Rapior S. 2023. Mushrooms as promising therapeutic resources: review and future perspectives. In Mushrooms with Therapeutic Potentials: Recent Advances in Research and Development, eds. Agrawal DC, Dhanasekeran M. Singapore: Springer. pp. 1–54 doi: 10.1007/978-981-19-9550-7_1

[37]

Badalyan S, Iotti M, Zambonelli A. 2026. The resource potential of mushrooms − a review. Mycological Spectrum 2(1):1−17

doi: 10.64993/MS.2.1.1
[38]

Badalyan SM, Zambonelli A. 2019. Biotechnological exploitation of macrofungi for the production of food, pharmaceuticals, and cosmeceuticals. In Advances in Macrofungi: Diversity, Ecology, and Biotechnology, eds. Sridhar KR, Deshmukh S. Boca Raton: CRC Press. pp. 199–230 doi: 10.1201/9780429504075

[39]

Badalyan SM, Zambonelli A. 2023. The potential of mushrooms in developing healthy food and biotech products. In Fungi and Fungal Products in Human Welfare and Biotechnology, eds. Satyanarayana T, Deshmukh SK. Singapore: Springer. pp. 307–344 doi: 10.1007/978-981-19-8853-0_11

[40]

González A, Cruz M, Losoya C, Nobre C, Loredo A, et al. 2020. Edible mushrooms as a novel protein source for functional foods. Food & Function 11(9):7400−7414

doi: 10.1039/D0FO01746A
[41]

Łysakowska P, Sobota A, Wirkijowska A. 2023. Medicinal mushrooms: their bioactive components, nutritional value, and application in functional food production − a review. Molecules 28(14):5393

doi: 10.3390/molecules28145393
[42]

Carvajal AESS, Koehnlein EA, Soares AA, Eler GJ, Nakashima ATA, et al. 2012. Bioactives of fruiting bodies and submerged culture mycelia of Agaricus brasiliensis (A. blazei) and their antioxidant properties. LWT 46(2):493−499

doi: 10.1016/j.lwt.2011.11.018
[43]

Elisashvili V. 2012. Submerged cultivation of medicinal mushrooms: bioprocesses and products (review). International Journal of Medicinal Mushrooms 14(3):211−239

doi: 10.1615/IntJMedMushr.v14.i3.10
[44]

Bakratsas G, Polydera A, Katapodis P, Stamatis H. 2021. Recent trends in submerged cultivation of mushrooms and their application as a source of nutraceuticals and food additives. Future Foods 4:100086

doi: 10.1016/j.fufo.2021.100086
[45]

Chang ST, Buswell JA. 1996. Mushroom nutriceuticals. World Journal of Microbiology and Biotechnology 12(5):473−476

doi: 10.1007/BF00419460
[46]

Badalyan SM, Barkhudaryan A, Rapior S. 2021. The cardioprotective properties of Agaricomycetes mushrooms growing in the territory of Armenia: review. International Journal of Medicinal Mushrooms 23(2):21−31

doi: 10.1615/IntJMedMushrooms.2021038280
[47]

Badalyan SM, Barkhudaryan A, Rapior S. 2022. Medicinal macrofungi as cosmeceuticals: a review. International Journal of Medicinal Mushrooms 24(4):1−13

doi: 10.1615/IntJMedMushrooms.2022043124
[48]

Rahi D, K Rahi, Chaudhary E. 2021. White-rot fungi in food and pharmaceutical industries. In Advances in Macrofungi: Industrial Avenues and Prospects, eds. Sridhar KR, Deshmukh SK. Boca Raton: CRC Press. pp. 175–206 doi: 10.1201/9781003191278-14

[49]

Sevindik M. 2021. Biological activities of Laetiporus species as a functional food. Journal of Mycology & Mycological Sciences 4(1):000138

doi: 10.23880/oajmms-16000138
[50]

Barua RC, Coniglio RO, Molina MA, Díaz GV, Fonseca MI. 2024. Fungi as biotechnological allies: exploring contributions of edible and medicinal mushrooms. Journal of Food Science 89:6888−6915

doi: 10.1111/1750-3841.17390
[51]

Takhtajyan AL. 1954. Map of the floristic regions of Armenian SSR. In Flora of Armenia. Vol 1. Yerevan: Academy of Sciences of the Armenian SSR. https://ace.aua.am/wp-content/uploads/2014/02/Takhtajyan-A.-L.-Flora-of-Armenia.-Volume-1.-Yerevan-Armenian-SSR-Academy-of-Science-Publishing-1954.pdf (in Russian)

[52]

Badalyan SM, Gharibyan NG, Gianchino C, Iotti M, Zambonelli A. 2023. Morphological observation and biomass formation in different edible medicinal Morchella collections (Pezizomycetes, Ascomycota). Italian Journal of Mycology 52:50−61

doi: 10.6092/issn.2531-7342/16112
[53]

Badalyan SM, Gharibyan NG. 2017. Characteristics of Mycelial Structures of Different Fungal Collections. Yerevan: YSU Press. http://publishing.ysu.am/hy/1505392977 (in Armenian/English)

[54]

Porter DL, Naleway SE. 2022. Hyphal systems and their effect on the mechanical properties of fungal sporocarps. Acta Biomaterialia 145:272−282

doi: 10.1016/j.actbio.2022.04.011
[55]

Clémençon H. 2004. Cytology and Plectology of the Hymenomycetes. Berlin/Stuttgart: J. Cramer. www.schweizerbart.de/publications/detail/isbn/9783443591014/Cytology_and_Plectology_of_the_Hymenomycetes#html

[56]

Neves MA. 1998. Estudos de culturas de fungos (Aphyllophorales) da Ilha de Santa Catarina, SC, Brasil. Biotemas 11:39−70

[57]

Motato-Vásquez V, Pires RM, Vitali VMV, de Mello Gugliotta A. 2016. Cultural and ligninolytic activity studies of some polypores (Basidiomycota) from Brazilian Atlantic Forest, São Paulo State, Brazil. Hoehnea 43(2):289−300

doi: 10.1590/2236-8906-81/2015
[58]

Saltarelli R, Ceccaroli P, Buffalini M, Vallorani L, Casadei L, et al. 2015. Biochemical characterization and antioxidant and antiproliferative activities of different Ganoderma collections. Journal of Molecular Microbiology and Biotechnology 25(1):16−25

doi: 10.1159/000369212
[59]

Badalyan SM, Gharibyan NG. 2016. Diversity of polypore bracket mushrooms, polyporales (Agaricomycetes), recorded in Armenia and their medicinal properties. International Journal of Medicinal Mushrooms 18:347−354

doi: 10.1615/IntJMedMushrooms.v18.i4.80
[60]

Adaskaveg JE, Gilbertson RL. 1989. Cultural studies of four North American species in the Ganoderma lucidum complex with comparisons to G. lucidum and G. tsugae. Mycological Research 92:182−191

doi: 10.1016/S0953-7562(89)80010-3
[61]

Moncalvo JM, Wang HF, Hseu RS. 1995. Gene phylogeny of the Ganoderma lucidum complex based on ribosomal DNA sequences. Comparison with traditional taxonomic characters. Mycological Research 99(12):1489−1499

doi: 10.1016/S0953-7562(09)80798-3
[62]

Kayashima T, Katayama T. 2002. Oxalic acid is available as a natural antioxidant in some systems. Biochimica et Biophysica Acta (BBA) − General Subjects 1573(1):1−3

doi: 10.1016/s0304-4165(02)00338-0
[63]

Connolly JH. 1997. Assessment of crystals for the identification of fungi. Northeastern Naturalist 4(4):279−284

doi: 10.2307/3858612
[64]

Xiao X, Huang T, Zhang J, Su Q, Tao L, et al. 2021. Evidence for the presence of hyphae and fruiting body calcium oxalate crystallites in Schizophyllum commune. bioRxiv Preprint:456293

doi: 10.1101/2021.08.13.456293
[65]

Krupodorova TA, Barshteyn VYu, Sekan AS. 2021. Review of the basic cultivation conditions influence on the growth of basidiomycetes. Current Research in Environmental & Applied Mycology 11(1):494−531

doi: 10.5943/cream/11/1/34