| [1] |
Schuurink R, Tissier A. 2020. Glandular trichomes: micro-organs with model status? |
| [2] |
Vimala, Hans D. 2024. Variations in epidermal trichomes of a mystic weed Parthenium hysterophorus L. from semi-arid regions of Barmer, Rajasthan (India). |
| [3] |
Antonious GF. 2001. Production and quantification of methyl ketones in wild tomato accessions. |
| [4] |
Ma L, Zheng SC, Zhang TK, Liu ZY, Wang XJ, et al. 2018. Effect of nicotine from tobacco root exudates on chemotaxis, growth, biocontrol efficiency, and colonization by Pseudomonas aeruginosa NXHG29. |
| [5] |
Molinaro F, Tyc O, Beekwilder J, Cankar K, Bertea CM, et al. 2018. The effect of isabelin, a sesquiterpene lactone from Ambrosia artemisiifolia on soil microorganisms and human pathogens. |
| [6] |
Jermakowicz E, Leśniewska J, Stocki M, Naczk AM, Kostro-Ambroziak A, et al. 2022. The floral signals of the inconspicuous orchid Malaxis monophyllos: how to lure small pollinators in an abundant environment. |
| [7] |
Moyano F, Cocucci A, Sérsic A. 2003. Accessory pollen adhesive from glandular trichomes on the anthers of Leonurus sibiricus L. (Lamiaceae). |
| [8] |
Yuan M, Sheng Y, Bao J, Wu W, Nie G, et al. 2025. AaMYC3 bridges the regulation of glandular trichome density and artemisinin biosynthesis in Artemisia annua. |
| [9] |
Hua B, Chang J, Wu M, Xu Z, Zhang F, et al. 2021. Mediation of JA signalling in glandular trichomes by the woolly/SlMYC1 regulatory module improves pest resistance in tomato. |
| [10] |
Feng Z, Bartholomew ES, Liu Z, Cui Y, Dong Y, et al. 2021. Glandular trichomes: new focus on horticultural crops. |
| [11] |
Karabourniotis G, Liakopoulos G, Nikolopoulos D, Bresta P. 2020. Protective and defensive roles of non-glandular trichomes against multiple stresses: structure–function coordination. |
| [12] |
Haratym W, Weryszko-Chmielewska E. 2017. Ultrastructural and histochemical analysis of glandular trichomes of Marrubium vulgare L. (Lamiaceae). |
| [13] |
Zhou Y, Tang N, Huang L, Zhao Y, Tang X, et al. 2018. Effects of salt stress on plant growth, antioxidant capacity, glandular trichome density, and volatile exudates of Schizonepeta tenuifolia Briq. |
| [14] |
Murungi LK, Kirwa H, Salifu D, Torto B. 2016. Opposing roles of foliar and glandular trichome volatile components in cultivated nightshade interaction with a specialist herbivore. |
| [15] |
Li Y, Yang Y, Li L, Tang K, Hao X, et al. 2024. Advanced metabolic engineering strategies for increasing artemisinin yield in Artemisia annua L. |
| [16] |
Turner GW, Gershenzon J, Croteau RB. 2000. Distribution of peltate glandular trichomes on developing leaves of peppermint. |
| [17] |
Wang J, Yuan S, Zhao Y, Shu X, Liu Z, et al. 2025. Wo interacts with SlTCP25 to regulate type I trichome branching in tomato. |
| [18] |
McDowell ET, Kapteyn J, Schmidt A, Li C, Kang JH, et al. 2011. Comparative functional genomic analysis of Solanum glandular trichome types. |
| [19] |
Zhang M, Li M, An Y, Liu C, Zhao Q, et al. 2025. Single-nucleus transcriptomics reveal the morphogenesis and artemisinin biosynthesis in Artemisia annua glandular trichomes. |
| [20] |
Lv D, Wang G, Zhang Q, Yu Y, Qin P-C, et al. 2022. Comparative transcriptome analysis of hard and tender fruit spines of cucumber to identify genes involved in the morphological development of fruit spines. |
| [21] |
Tian Z, Luo Q, Zuo Z. 2021. Seasonal emission of monoterpenes from four chemotypes of Cinnamomum camphora. |
| [22] |
Kamatou GP, Vermaak I, Viljoen AM, Lawrence BM. 2013. Menthol: a simple monoterpene with remarkable biological properties. |
| [23] |
Froldi G, Benetti F, Mondin A, Roverso M, Pangrazzi E, et al. 2023. Pterodon emarginatus seed preparations: antiradical activity, chemical characterization, and in silico ADMET parameters of β-caryophyllene and farnesol. |
| [24] |
Hébert M, Bellavance G, Barriault L. 2025. Advanced synthetic strategies toward ginkgolide diterpenoids: total synthesis of (±)-ginkgolide C and formal syntheses of (±)-ginkgolides A and B. |
| [25] |
Xu W, Zhang JH, Liu X, Jia AL, Liu XL, et al. 2017. Two new dammarane-type triterpenoid saponins from ginseng medicinal fungal substance. |
| [26] |
Ravichandran B, Sanniyasi E, Kuppusamy SG, Perumal P. 2024. Onco-therapeutic effect of novel triterpenoid compound oleanolic acid isolated and characterized from the methanolic extract of Coldenia procumbens (Linn.). |
| [27] |
Frey M, Jochimsen CM, Degenhardt J. 2025. Cytochrome P450-induced backbone rearrangements in terpene biosynthesis of plants. |
| [28] |
Ma Y, Chen Q, Wang Y, Zhang F, Wang C, Wang G. 2023. Heteromerization of short-chain trans-prenyltransferase controls precursor allocation within a plastidial terpenoid network. |
| [29] |
Bergman ME, Phillips MA. 2021. Structural diversity and biosynthesis of plant derived p-menthane monoterpenes. |
| [30] |
Salehi M, Karimzadeh G, Naghavi MR, Naghdi Badi H, Rashidi Monfared S. 2018. Expression of key genes affecting artemisinin content in five Artemisia species. |
| [31] |
Sallaud C, Giacalone C, Töpfer R, Goepfert S, Bakaher N, et al. 2012. Characterization of two genes for the biosynthesis of the labdane diterpene Z-abienol in tobacco (Nicotiana tabacum) glandular trichomes. |
| [32] |
Booth JK, Yuen MMS, Jancsik S, Madilao LL, Page JE, Bohlmann J. 2020. Terpene synthases and terpene variation in Cannabis sativa. |
| [33] |
Patzak J, Henychová A, Matoušek J. 2021. Developmental regulation of lupulin gland-associated genes in aromatic and bitter hops (Humulus lupulus L.). |
| [34] |
Turner GW, Gershenzon J, Croteau RB. 2000. Development of peltate glandular trichomes of peppermint. |
| [35] |
Huchelmann A, Boutry M, Hachez C. 2017. Plant glandular trichomes: natural cell factories of high biotechnological interest. |
| [36] |
Bergau N, Bennewitz S, Syrowatka F, Hause G, Tissier A. 2015. The development of type VI glandular trichomes in the cultivated tomato Solanum lycopersicum and a related wild species S. habrochaites. |
| [37] |
Gang DR, Simon J, Lewinsohn E, Pichersky E. 2002. Peltate glandular trichomes of Ocimum basilicum L. (sweet basil) contain high levels of enzymes involved in the biosynthesis of phenylpropenes. In Breeding Research on Aromatic and Medicinal Plants, eds. Johnson CB, Franz C. Boca Raton: CRC Press. pp. 189−195 doi: 10.1201/9781003573227-28 |
| [38] |
Zhang Y, Wang D, Li H, Bai H, Sun M, et al. 2023. Formation mechanism of glandular trichomes involved in the synthesis and storage of terpenoids in lavender. |
| [39] |
Giuliani C, Ascrizzi R, Corrà S, Bini LM, Flamini G, et al. 2017. Ultrastructural insight into terpene-producing trichomes and essential oil profile in Salvia greggii A. gray. |
| [40] |
Gersbach PV. 2002. The essential oil secretory structures of Prostanthera ovalifolia (Lamiaceae). |
| [41] |
Balcke GU, Bennewitz S, Bergau N, Athmer B, Henning A, et al. 2017. Multi-omics of tomato glandular trichomes reveals distinct features of central carbon metabolism supporting high productivity of specialized metabolites. |
| [42] |
Sallets A, Beyaert M, Boutry M, Champagne A. 2014. Comparative proteomics of short and tall glandular trichomes of Nicotiana tabacum reveals differential metabolic activities. |
| [43] |
Holopainen JK, Gershenzon J. 2010. Multiple stress factors and the emission of plant VOCs. |
| [44] |
Xie L, Yan T, Li L, Chen M, Ma Y, et al. 2021. The WRKY transcription factor AaGSW2 promotes glandular trichome initiation in Artemisia annua. |
| [45] |
Hu X, Liu H, Chen T, Ye Y, Shao J, et al. 2025. AaPDF2 promotes glandular trichome formation in Artemisia annua through jasmonic acid signaling. |
| [46] |
Yan T, Li L, Xie L, Chen M, Shen Q, et al. 2018. A novel HD-ZIP IV/MIXTA complex promotes glandular trichome initiation and cuticle development in Artemisia annua. |
| [47] |
Shi P, Fu X, Shen Q, Liu M, Pan Q, et al. 2018. The roles of AaMIXTA1 in regulating the initiation of glandular trichomes and cuticle biosynthesis in Artemisia annua. |
| [48] |
Gao Y, Ruan J, Wang G, Zhang Y, Wang Y, et al. 2025. SlHDZIV8-SlnsLTP33 module controls trichome density and lipid metabolism in tomato. |
| [49] |
Gao S, Gao Y, Xiong C, Yu G, Chang J, et al. 2017. The tomato B-type cyclin gene, SlCycB2, plays key roles in reproductive organ development, trichome initiation, terpenoids biosynthesis and Prodenia litura defense. |
| [50] |
Zhang H, Xu H, Xu M, Yan X, Wang Z, et al. 2023. Transcription factors NtHD9 and NtHD12 control long glandular trichome formation via jasmonate signaling. |
| [51] |
Tian Y, Dai TW, Fan CX, Zhou J, Ren XI, et al. 2020. An HD-Zip IV transcription factor protein NbGL3 regulates glandular trichome initiation in tobacco. |
| [52] |
Khan RA, Kumar A, Abbas N. 2024. A bHLH transcription factor AaMYC2-type positively regulates glandular trichome density and artemisinin biosynthesis in Artemisia annua. |
| [53] |
Fu X, Peng B, Hassani D, Xie L, Liu H, et al. 2021. AaWRKY9 contributes to light- and jasmonate-mediated to regulate the biosynthesis of artemisinin in Artemisia annua. |
| [54] |
Lv Z, Li J, Qiu S, Qi F, Su H, et al. 2022. The transcription factors TLR1 and TLR2 negatively regulate trichome density and artemisinin levels in Artemisia annua. |
| [55] |
Yu L, Zhang Y, Ding Q, Wang H, Meng X, et al. 2024. The SlMYC1-TOR module regulates trichome formation and terpene biosynthesis in tomatoes (Solanum lycopersicum L.). |
| [56] |
Xu J, van Herwijnen ZO, Dräger DB, Sui C, Haring MA, et al. 2018. SlMYC1 regulates type VI glandular trichome formation and terpene biosynthesis in tomato glandular cells. |
| [57] |
Gong Z, Luo Y, Zhang W, Jian W, Zhang L, et al. 2021. A SlMYB75-centred transcriptional cascade regulates trichome formation and sesquiterpene accumulation in tomato. |
| [58] |
Guan Y, Jiang L, Wang Y, Liu G, Wu J, et al. 2024. CmMYC2-CmMYBML1 module orchestrates the resistance to herbivory by synchronously regulating the trichome development and constitutive terpene biosynthesis in Chrysanthemum morifolium. |
| [59] |
Zhan J, Di T, Chen X, Zheng T, Sun W, et al. 2024. CbMYB108 integrates the regulation of diterpene biosynthesis and trichome development in Conyza blinii against UV-B. |
| [60] |
Chen Q, Li L, Qi X, Fang H, Yu X, et al. 2023. The non-specific lipid transfer protein McLTPII. 9 of Mentha canadensis is involved in peltate glandular trichome density and volatile compound metabolism. |
| [61] |
Shi J, Zhang J, Sun L, Tang X, Wang K. 2026. Jasmonate-responsive transcription factor StMYC2 coordinately regulates peltate glandular trichome development and monoterpenoid biosynthesis in Schizonepeta tenuifolia. |
| [62] |
Yu X, Liu Z, Sun X. 2023. Single-cell and spatial multi-omics in the plant sciences: technical advances, applications, and perspectives. |
| [63] |
Wang K, Liu Z. 2025. Plant synthetic biology-based biofortification, strategies and recent progresses. |
| [64] |
Yang JS, Reyna-Llorens I. 2023. Plant synthetic biology: exploring the frontiers of sustainable agriculture and fundamental plant biology. |
| [65] |
Sun X, Wu X, Jiang X, Huang H, Yang A, et al. 2026. Heterologous production of forskolin in tobacco (Nicotiana tabacum) via glandular trichome specific engineering and metabolic flux redirection. |