[1]

Wang R, Yang Y, Wang X, Li J, Gao Y, et al. 2025. Response of seed germination and seedling growth of perennial ryegrass (Lolium perenne L.) to drought, salinity, and pH in Karst regions. Scientific Reports 15:16874

doi: 10.1038/s41598-025-01539-5
[2]

Chen S, Xie Y, Pan S, Yu S, Zhang L. 2025. Identification of perennial ryegrass CDPK gene family and function exploration of LpCDPK27 upon salt stress. Grass Research 5:e009

doi: 10.48130/grares-0025-0010
[3]

Han K, Zhao Y, Sun YH, Li Y. 2023. NACs, generalist in plant life. Plant Biotechnology Journal 21:2433−2457

doi: 10.1111/pbi.14161
[4]

Chen Y, Xia P. 2025. NAC transcription factors as biological macromolecules responded to abiotic stress: a comprehensive review. International Journal of Biological Macromolecules 308:142400

doi: 10.1016/j.ijbiomac.2025.142400
[5]

Mao H, Wang H, Liu S, Li Z, Yang X, et al. 2015. A transposable element in a NAC gene is associated with drought tolerance in maize seedlings. Nature Communications 6:8326

doi: 10.1038/ncomms9326
[6]

Mao H, Li S, Chen B, Jian C, Mei F, et al. 2022. Variation in cis-regulation of a NAC transcription factor contributes to drought tolerance in wheat. Molecular Plant 15:276−292

doi: 10.1016/j.molp.2021.11.007
[7]

Yu X, Xie Y, Wang L, Li L, Jiang S, et al. 2024. Transcription factor NAC78 cooperates with NAC78 interacting protein 6 to confer drought tolerance in rice. Plant Physiology 196:1642−1658

doi: 10.1093/plphys/kiae395
[8]

Duan M, Zhang R, Zhu F, Zhang Z, Gou L, et al. 2017. A lipid-anchored NAC transcription factor is translocated into the nucleus and activates Glyoxalase I expression during drought stress. The Plant Cell 29:1748−1772

doi: 10.1105/tpc.17.00044
[9]

Hao H, Zhou Q, Yin T, Dong C, Zhang Z, et al. 2026. LpCbDR1 regulates leaf senescence and drought tolerance by activating the chlorophyll b reductase gene and stress-related genes in perennial ryegrass. Horticulture Research 13(7):uhag093

doi: 10.1093/hr/uhag093
[10]

Qiang Z, Zeng Z, Ma D, Li J, Zhao Y, et al. 2025. NAC transcription factor LpNAC22 positively regulates drought tolerance in perennial ryegrass. Plant, Cell & Environment 48:7256−7270

doi: 10.1111/pce.70022
[11]

Zhang WJ, Dewey RE, Boss W, Phillippy BQ, Qu R. 2013. Enhanced Agrobacterium-mediated transformation efficiencies in monocot cells is associated with attenuated defense responses. Plant Molecular Biology 81:273−286

doi: 10.1007/s11103-012-9997-8
[12]

Sun T, Wang W, Hu X, Meng L, Xiang L, et al. 2024. HSFA3 functions as a positive regulator of HSFA2a to enhance thermotolerance in perennial ryegrass. Plant Physiology and Biochemistry 208:108512

doi: 10.1016/j.plaphy.2024.108512
[13]

She M, Zheng D, Zhang S, Ke Z, Wu Z, et al. 2024. Functional analysis of maize GRAS transcription factor gene ZmGRAS72 in response to drought and salt stresses. Agriculture Communications 2:100054

doi: 10.1016/j.agrcom.2024.100054
[14]

Qiang Z, Sun H, Ge F, Li W, Li C, et al. 2022. The transcription factor ZmMYB69 represses lignin biosynthesis by activating ZmMYB31/42 expression in maize. Plant Physiology 189:1916−1919

doi: 10.1093/plphys/kiac233
[15]

Wang C, Liu S, Dong Y, Zhao Y, Geng A, et al. 2016. PdEPF1 regulates water-use efficiency and drought tolerance by modulating stomatal density in poplar. Plant Biotechnology Journal 14:849−860

doi: 10.1111/pbi.12434
[16]

Zhang Y, Yu S, Niu P, Su L, Jiao X, et al. 2024. RcMYB8 enhances salt and drought tolerance in rose (Rosa chinensis) by modulating RcPR5/1 and RcP5CS1. Molecular Horticulture 4:3

doi: 10.1186/s43897-024-00080-9
[17]

Ding X, Jiang Y, Zhao H, Guo D, He L, et al. 2018. Electrical conductivity of nutrient solution influenced photosynthesis, quality, and antioxidant enzyme activity of pakchoi (Brassica campestris L. ssp. Chinensis) in a hydroponic system. PLoS One 13:e0202090

doi: 10.1371/journal.pone.0202090
[18]

Guo H, Wang Y, Wang L, Hu P, Wang Y, et al. 2017. Expression of the MYB transcription factor gene BplMYB46 affects abiotic stress tolerance and secondary cell wall deposition in Betula platyphylla. Plant Biotechnology Journal 15:107−121

doi: 10.1111/pbi.12595
[19]

Zhang X, Wang L, Meng H, Wen H, Fan Y, et al. 2011. Maize ABP9 enhances tolerance to multiple stresses in transgenic Arabidopsis by modulating ABA signaling and cellular levels of reactive oxygen species. Plant Molecular Biology 75:365−378

doi: 10.1007/s11103-011-9732-x
[20]

Shi J, Fu XZ, Peng T, Huang XS, Fan QJ, et al. 2010. Spermine pretreatment confers dehydration tolerance of citrus in vitro plants via modulation of antioxidative capacity and stomatal response. Tree Physiology 30:914−922

doi: 10.1093/treephys/tpq030
[21]

Livak KJ, Schmittgen TD. 2001. Analysis of relative gene expression data using real-time quantitative PCR and the 2−ΔΔCᴛ method. Methods 25:402−408

doi: 10.1006/meth.2001.1262
[22]

Ghosh UK, Islam MN, Siddiqui MN, Cao X, Khan MAR. 2022. Proline, a multifaceted signalling molecule in plant responses to abiotic stress: understanding the physiological mechanisms. Plant Biology 24:227−239

doi: 10.1111/plb.13363
[23]

Xiong H, He H, Chang Y, Miao B, Liu Z, et al. 2025. Multiple roles of NAC transcription factors in plant development and stress responses. Journal of Integrative Plant Biology 67:510−538

doi: 10.1111/jipb.13854
[24]

Ernst HA, Olsen AN, Skriver K, Larsen S, Lo Leggio L. 2004. Structure of the conserved domain of ANAC, a member of the NAC family of transcription factors. EMBO Reports 5:19

doi: 10.1038/sj.embor.7400093
[25]

Jensen MK, Skriver K. 2014. NAC transcription factor gene regulatory and protein-protein interaction networks in plant stress responses and senescence. IUBMB Life 66:156−166

doi: 10.1002/iub.1256
[26]

Mohanty S, Hembram P. 2025. An overview of LEA genes and their importance in combating abiotic stress in rice. Plant Molecular Biology Reporter 43:337−351

doi: 10.1007/s11105-024-01468-z
[27]

Liu F, Zhao Y, Wang X, Wang B, Xiao F, et al. 2023. Physiological response and drought resistance evaluation of Gleditsia sinensis seedlings under drought-rehydration state. Scientific Reports 13:19963

doi: 10.1038/s41598-023-45394-8
[28]

Ashraf M, Foolad MR. 2007. Roles of glycine betaine and proline in improving plant abiotic stress resistance. Environmental and Experimental Botany 59:206−216

doi: 10.1016/j.envexpbot.2005.12.006
[29]

Thirumalaikumar VP, Devkar V, Mehterov N, Ali S, Ozgur R, et al. 2018. NAC transcription factor JUNGBRUNNEN1 enhances drought tolerance in tomato. Plant Biotechnology Journal 16:354−366

doi: 10.1111/pbi.12776
[30]

Lu J, Wang W, Yang S, Shi L, Song F, et al. 2025. Transcription factor OsNAC29a confers drought tolerance through the ABA pathway in rice. Plant Physiology and Biochemistry 225:109989

doi: 10.1016/j.plaphy.2025.109989
[31]

Zhao JL, Wu Q, Wu HL, Wang AH, Wang XL, et al. 2022. FtNAC31, a Tartary buckwheat NAC transcription factor, enhances salt and drought tolerance in transgenic Arabidopsis. Plant Physiology and Biochemistry 191:20−33

doi: 10.1016/j.plaphy.2022.09.016