[1]

Sang J, Zhang Y, Sang J, Li CQ. 2019. Anthocyanins from Nitraria tangutorun: qualitative and quantitative analyses, antioxidant and anti-inflammatory activities and their stabilities as affected by some phenolic acids. Journal of Food Measurement and Characterization 13:421−430

doi: 10.1007/s11694-018-9956-4
[2]

Li X, Liu H, Li C, Li Y. 2024. A systematic review on the morphology structure, propagation characteristics, resistance physiology and exploitation and utilization of Nitraria tangutorum Bobrov. PeerJ 12:e17830

doi: 10.7717/peerj.17830
[3]

Ren S, Lv G, Teng D. 2021. Anatomical structure of Nitraria spp. leaves from different habitats in Southern Xinjiang, China. Human and Ecological Risk Assessment: an International Journal 27:790−803

doi: 10.1080/10807039.2020.1761244
[4]

Chen S, Zhou H, Zhang G, Dong Q, Wang Z, et al. 2021. Characterization, antioxidant, and neuroprotective effects of anthocyanins from Nitraria tangutorum Bobr. fruit. Food Chemistry 353:129435

doi: 10.1016/j.foodchem.2021.129435
[5]

Bao X, Zong Y, Hu N, Li S, Liu B, et al. 2022. Functional R2R3-MYB transcription factor NsMYB1, regulating anthocyanin biosynthesis, was relative to the fruit color differentiation in Nitraria sibirica Pall. BMC Plant Biology 22:186

doi: 10.1186/s12870-022-03561-5
[6]

Ren L, Dong Q, Liu Z, Wang Y, Tan N, et al. 2024. Optimization of subcritical water extraction, UPLC-triple-TOF-MS/MS analysis, antioxidant and α-glucosidase inhibitory activities of anthocyanins from Nitraria sibirica Pall. fruits. Food Chemistry 23:101626

doi: 10.1016/j.fochx.2024.101626
[7]

Chang Y, Lv G, He X. 2024. Growth adaptation of Nitraria sibirica in drought: Hormone-mediated flavonoid accumulation, altered metabolic pathways, and decreased network complexity. Environmental and Experimental Botany 221:105720

doi: 10.1016/j.envexpbot.2024.105720
[8]

Hadj Salem J, Chevalot I, Harscoat-Schiavo C, Paris C, Fick M, et al. 2011. Biological activities of flavonoids from Nitraria retusa (Forssk.) Asch. and their acylated derivatives. Food Chemistry 124:486−494

doi: 10.1016/j.foodchem.2010.06.059
[9]

Zhao Q, Zhang J, Li Y, Yang Z, Wang Q, et al. 2024. Integrated metabolomic and transcriptomic analysis of Nitraria berries indicate the role of flavonoids in adaptation to high altitude. Metabolites 14:591

doi: 10.3390/metabo14110591
[10]

Yue H, Li J, Wu L, Wang F, Li Z, et al. 2025. Tentative characterization of three novel coumaroyl glucoside β-carboline alkaloids and discovery of hypoglycemic compounds from Nitraria tangutorum Bobr. fruit. Food Chemistry 481:143923

doi: 10.1016/j.foodchem.2025.143923
[11]

Wu D, Jiang S, Wang G, Wang L, Wu L, et al. 2024. Characterization of alkaloids and phenolics in Nitraria roborowskii Kom. fruit by UHPLC-triple-TOF-MS/MS and its sucrase and maltase inhibitory effects. Food Chemistry 447:138743

doi: 10.1016/j.foodchem.2024.138743
[12]

Jiang S, Chen C, Dong Q, Shao Y, Zhao X, et al. 2021. Alkaloids and phenolics identification in fruit of Nitraria tangutorum Bobr. by UPLC-Q-TOF-MS/MS and their a-glucosidase inhibitory effects in vivo and in vitro. Food Chemistry 364:130412

doi: 10.1016/j.foodchem.2021.130412
[13]

Meng J, Deng K, Hu N, Wang H. 2021. Nitraria tangutorum Bobr.-derived polysaccharides protect against LPS-induced lung injury. International Journal of Biological Macromolecules 186:71−78

doi: 10.1016/j.ijbiomac.2021.06.181
[14]

Abuduwaili A, Mutailifu P, Nuerxiati R, Gao Y, Aisa HA, et al. 2021. Structure and biological activity of polysaccharides from Nitraria sibirica pall fruit. Food Bioscience 40:100903

doi: 10.1016/j.fbio.2021.100903
[15]

Rjeibi I, Feriani A, Hentati F, Hfaiedh N, Michaud P, et al. 2019. Structural characterization of water-soluble polysaccharides from Nitraria retusa fruits and their antioxidant and hypolipidemic activities. International Journal of Biological Macromolecules 129:422−432

doi: 10.1016/j.ijbiomac.2019.02.049
[16]

Zar Kalai F, Dakhlaoui S, Hammami M, Mkadmini K, Ksouri R, et al. 2022. Phenolic compounds and biological activities of different organs from aerial part of Nitraria retusa (Forssk.) Asch.: effects of solvents. International Journal of Food Properties 25:1524−1538

doi: 10.1080/10942912.2022.2087673
[17]

Turghun C, Bakri M, Abdulla R, Ma Q, Aisa HA. 2020. Comprehensive characterisation of phenolics from Nitraria sibirica leaf extracts by UHPLC-quadrupole-orbitrap-MS and evaluation of their anti-hypertensive activity. Journal of Ethnopharmacology 261:113019

doi: 10.1016/j.jep.2020.113019
[18]

Almasarwah SY, Oran SA, Darwish RM. 2023. Efficacy of Nitraria retusa L. fruits aqueous and methanol extracts as antioxidant and anti-inflammatory activities on carrageenan-induced paw edema in rats. Tropical Journal of Natural Product Research 7:2725−2729

doi: 10.26538/tjnpr/v7i4.9
[19]

Boubaker J, Lahmar A, Salek A, Kriffa M, Chekir-Ghedira L. 2025. Cytotoxic effect of antioxidant products from Nitraria retusa leaves in combination with temozolomide on glioblastoma cell growth. Research Square Preprint

doi: 10.21203/rs.3.rs-5687018/v1
[20]

Rjeibi I, Hentati F, Feriani A, Hfaiedh N, Delattre C, et al. 2019. Novel antioxidant, anti-α-amylase, anti-inflammatory and antinociceptive water-soluble polysaccharides from the aerial part of Nitraria retusa. Foods 9:28

doi: 10.3390/foods9010028
[21]

Mariem C, Sameh M, Nadhem S, Soumaya Z, Najiba Z, et al. 2014. Antioxidant and antimicrobial properties of the extracts from Nitraria retusa fruits and their applications to meat product preservation. Industrial Crops and Products 55:295−303

doi: 10.1016/j.indcrop.2014.01.036
[22]

Chen GL, Qi J, Zhan G, Jun R, Ren L. 2015. Antioxidant and antimicrobial activities of extracts from fruits of Nitraria sibirica Pall. in vitro. Acta Horticulturae 1106:137−142

doi: 10.17660/ActaHortic.2015.1106.21
[23]

Mohamed AA, Ali SI, Darwesh OM, El-Hallouty SM, Sameeh MY. 2015. Chemical compositions, potential cytotoxic and antimicrobial activities of Nitraria retusa methanolic extract sub-fractions. International Journal of Toxicology and Pharmacology Research 7:204−212

[24]

Jiang S, Wang L, Jia W, Wu D, Wu L, et al. 2023. Hypoglycemic effect of Nitraria tangutorum fruit by inhibiting glycosidase and regulating IRS1/PI3K/AKT signalling pathway and its active ingredient identification by UPLC-MS. Food & Function 14:7869−7881

doi: 10.1039/d3fo02495d
[25]

Jia QQ, Yang ZF, Wang Q, Zhao Q, Jia YJ, et al. 2023. Chemical profiling of Nitraria roborowskii Kom. by UPLC-Q-Orbitrap-MS and their hypolipidemic effects in vivo. Chemistry & Biodiversity 20:e202300683

doi: 10.1002/cbdv.202300683
[26]

Laouani A, Nasrallah H, Sassi A, Ferdousi F, Kalai FZ, et al. 2024. Antiobesity and hypolipidemic potential of Nitraria retusa extract in overweight/obese women: a randomized, double-blind, placebo-controlled pilot study. Nutrients 16:317

doi: 10.3390/nu16020317
[27]

Dong Q, Hu N, Suo Y, Chi X, Wang H. 2019. The complete chloroplast genome sequences of two species from Nitraria. Mitochondrial DNA Part B 4:1229−1230

doi: 10.1080/23802359.2019.1591225
[28]

Song F, Feng Y. 2021. The complete chloroplast genome of the desert shrub Nitraria sphaerocarpa (Nitrariaceae) and phylogenetic analysis. Mitochondrial DNA Part B 6:1901−1903

doi: 10.1080/23802359.2021.1934580
[29]

Ma X, Ru D, Morales-Briones DF, Mei F, Wu J, et al. 2023. Genome sequence and salinity adaptation of the desert shrub Nitraria sibirica (Nitrariaceae, Sapindales). DNA Research 30:dsad011

doi: 10.1093/dnares/dsad011
[30]

Wu X, Zhu J, Zhu L, Tang Y, Hao Z, et al. 2023. Genome-wide analyses of calmodulin and calmodulin-like proteins in the halophyte Nitraria sibirica reveal their involvement in response to salinity, drought and cold stress. International Journal of Biological Macromolecules 253:127442

doi: 10.1016/j.ijbiomac.2023.127442
[31]

Wang J, Dang Z, Zhang H, Zheng L, Borjigin T, et al. 2016. Gene transcript profiles in the desert plant Nitraria tangutorum during fruit development and ripening. Molecular Genetics and Genomics 291:383−398

doi: 10.1007/s00438-015-1116-5
[32]

Li Y, Wang S, Zhao Q, Wang Q, Yang Z, et al. 2025. Integrating metabolomics and transcriptomics comprehensively reveals the global metabolic differences in three species of Nitraria berries. Scientific Reports 15:15507

doi: 10.1038/s41598-025-00445-0
[33]

Li H, Tang X, Yang X, Zhang H. 2021. Comprehensive transcriptome and metabolome profiling reveal metabolic mechanisms of Nitraria sibirica Pall. to salt stress. Scientific Reports 11:12878

doi: 10.1038/s41598-021-92317-6
[34]

Tang X, Wang R, Yang X, Zhu J, Liu Z. et al. 2014. Isolation and expression analysis of a vacuolar membrane Na+/H+ antiporter gene NtNHX1 from Nitraria tangutorum. Scientia Silvae Sinicae 50:38−44 (in Chinese)

[35]

Tang X, Zhang H, Shabala S, Li H, Yang X, et al. 2021. Tissue tolerance mechanisms conferring salinity tolerance in a halophytic perennial species Nitraria sibirica Pall. Tree Physiology 41:1264−1277

doi: 10.1093/treephys/tpaa174
[36]

Lu L, Wang Y, Chen Y, Zhu L, Wu X, et al. 2025. Salt stimulates carbon fixation in the halophyte Nitraria sibirica to enhance growth. Forestry Research 5:e004

doi: 10.48130/forres-0025-0004
[37]

Zhang N, Cahaeraduqin S, Du J, Pan Y, Wang R, et al. 2024. Effects of exogenous H2O2 on ROS metabolism in Nitraria tangutorum Bobr. leaves under salt stress. Journal of Northeast Agricultural University (English Edition) 31:20−27

[38]

Zheng L, Dang Z, Li H, Zhang H, Wu S, et al. 2014. Isolation and characterization of a Δ1-pyrroline-5-carboxylate synthetase (NtP5CS) from Nitraria tangutorum Bobr. and functional comparison with its Arabidopsis homologue. Molecular Biology Reports 41:563−572

doi: 10.1007/s11033-013-2893-8
[39]

Didi DA, Su S, Sam FE, Tiika RJ, Zhang X. 2022. Effect of plant growth regulators on osmotic regulatory substances and antioxidant enzyme activity of Nitraria tangutorum. Plants 11:2559

doi: 10.3390/plants11192559
[40]

Li X, Liu H, He C, Li Y. 2025. Physiological mechanisms of exogenous ABA in alleviating drought stress in Nitraria tangutorum. Plants 14:2643

doi: 10.3390/plants14172643
[41]

Gao Z, Gao S, Li P, Zhang Y, Ma B, et al. 2021. Exogenous methyl jasmonate promotes salt stress-induced growth inhibition and prioritizes defense response of Nitraria tangutorum Bobr. Physiologia Plantarum 172:162−175

doi: 10.1111/ppl.13314
[42]

Zhang J, Cheng K, Liu X, Dai Z, Zheng L, et al. 2023. Exogenous abscisic acid and sodium nitroprusside regulate flavonoid biosynthesis and photosynthesis of Nitraria tangutorum Bobr in alkali stress. Frontiers in Plant Science 14:1118984

doi: 10.3389/fpls.2023.1118984
[43]

Liu C, Duan N, Chen X, Li H, Zhao X, et al. 2022. Metabolic pathways involved in the drought stress response of Nitraria tangutorum as revealed by transcriptome analysis. Forests 13:509

doi: 10.3390/f13040509
[44]

Wang YM, Yu CH, Zhao XJ, Zhao JQ. 2020. A rapid high-performance liquid chromatography separation of a new anthocyanin from Nitraria tangutorum. Journal of Asian Natural Products Research 22:503−507

doi: 10.1080/10286020.2019.1593968
[45]

Zhang M, Ma J, Bi H, Song J, Yang H, et al. 2017. Characterization and cardioprotective activity of anthocyanins from Nitraria tangutorum Bobr. by-products. Food & Function 8:2771−2782

doi: 10.1039/c7fo00569e
[46]

Zheng J, Li H, Ding C, Suo Y, Wang L, et al. 2011. Anthocyanins composition and antioxidant activity of two major wild Nitraria tangutorun Bobr. variations from Qinghai–Tibet Plateau. Food Research International 44:2041−2046

doi: 10.1016/j.foodres.2010.07.008
[47]

Gao Z, Wang YC, Chang YX. 2016. Determination of flavonoids and anthocyanins in Nitraria tangutorum by high performance liquid chromatography coupled with tandem mass spectrometry. Protein & Peptide Letters 23:424−432

doi: 10.2174/0929866523666160314152447
[48]

Zou Q, Feng J, Li T, Cheng G, Wang W, et al. 2022. Antioxidation and anti-inflammatory actions of the extract of Nitraria Tangutorum Bobr. fruits reduce the severity of ulcerative colitis in a dextran sulphate sodium-induced mice model. Journal of Functional Foods 91:105005

doi: 10.1016/j.jff.2022.105005
[49]

Halim AF, Saad HEA, Hashish NE. 1995. Flavonol glycosides from Nitraria retusa. Phytochemistry 40:349−351

doi: 10.1016/0031-9422(95)00167-6
[50]

Jiang S, Zhang Y, Zhao X, Shao Y, Wei W, et al. 2021. A new flavonol acylglycoside from the fruits of Nitraria tangutorum Bobr. Natural Product Research 35:3652−3657

doi: 10.1080/14786419.2020.1721487
[51]

Boubaker J, Ben Sghaier M, Skandrani I, Ghedira K, Chekir-Ghedira L. 2012. Isorhamnetin 3-O-robinobioside from Nitraria retusa leaves enhance antioxidant and antigenotoxic activity in human chronic myelogenous leukemia cell line K562. BMC Complementary and Alternative Medicine 12:135

doi: 10.1186/1472-6882-12-135
[52]

Kalai FZ, Boulaaba M, Ksouri R, Isoda H. 2021. Identification of main phenolic compounds of Nitraria retusa leaf and stem extracts and evaluation of their anti-adipogenic activity in 3T3-L1 cells. Research Square Preprint

doi: 10.21203/rs.3.rs-868059/v1
[53]

Boubaker J, Lahmar A, Salek A, Selmi M, Kriffa M, et al. 2025. Antitumoral effect of Nitraria retusa bioactive compounds on two glioblastoma cell lines. Human Cell 39:23

doi: 10.1007/s13577-025-01334-4
[54]

Jia Q, Zhang S, Zhang H, Yang X, Cui X, et al. 2020. A comparative study on polyphenolic composition of berries from the Tibetan Plateau by UPLC-Q-Orbitrap MS system. Chemistry & Biodiversity 17:e2000033

doi: 10.1002/cbdv.202000033
[55]

Khowdiary MM, Alatawi Z, Alhowiti A, Amin MA, Daghistani H, et al. 2024. Phytochemical analysis and multifaceted biomedical activities of Nitraria retusa extract as natural product-based therapies. Life 14:1629

doi: 10.3390/life14121629
[56]

Song Q, Zhao H, Fu Z, Chen D, Lu Y. 2018. Antioxidant and anticomplement compounds isolated from Nitraria sibirica fruit by high-speed counter-current chromatography. Pharmacognosy Magazine 14:541−547

doi: 10.4103/pm.pm_207_18
[57]

Turghun C, Bakri M, Liu GY, Bobakulov K, Aisa HA. 2021. Phenolic glycosides from Nitraria sibirica leaves and their in vitro biological activities. Natural Product Research 35:1388−1392

doi: 10.1080/14786419.2019.1647429
[58]

Song Q, Xia X, Ji C, Chen D, Lu Y. 2019. Optimized flash extraction and UPLC-MS analysis on antioxidant compositions of Nitraria sibirica fruit. Journal of Pharmaceutical and Biomedical Analysis 172:379−387

doi: 10.1016/j.jpba.2019.05.014
[59]

Senejoux F, Girard C, Aisa HA, Bakri M, Kerram P, et al. 2012. Vasorelaxant and hypotensive effects of a hydroalcoholic extract from the fruits of Nitraria sibirica Pall. (Nitrariaceae). Journal of Ethnopharmacology 141:629−634

doi: 10.1016/j.jep.2011.08.012
[60]

Kalai FZ, Mkadmini K, Hammami M, Isoda H, Ksouri R. 2022. Intensification of phenolic compounds extraction from Nitraria retusa leaves by ultrasound-assisted system using box–behnken design and evaluation of biological activities. Biological and Chemical Research 9:19−34

[61]

Rafiee D, Ebrahimi MA, Qavami N, Zarinpanjeh N. 2023. The effect of NaCl and salicylic acid on total phenolic and flavonoid contents in suspension culture of Nitraria schoberi. Journal of Medicinal plants and By-products 12:397−404

doi: 10.22092/jmpb.2022.358302.1465
[62]

Yang IF, Liub C. 2024. Combined microbial fermentation converts bioactive compounds in Nitraria tangutorum Bobrov fruit and displays its antidiabetic potential. JSM Biotechnology and Biomedical Engineering 9:1093

doi: 10.47739/2333-7117.biotechnology.1093
[63]

Tulyaganov TS, Nazarov OM, Levkovich MG, Abdullaev ND. 2001. Alkaloids of the Nitraria genus. komavine and acetylkomavine. Chemistry of Natural Compounds 37:61−64

doi: 10.1023/A:1017606727632
[64]

Tulyaganov TS, Nazarov OM. 2000. Alkaloids of Nitraria schoberi. N-methylnitrarine. Chemistry of Natural Compounds 36:393−395

doi: 10.1023/A:1002853132702
[65]

Zhao JQ, Wang YM, Yang YL, Zeng Y, Wang QL, et al. 2017. Isolation and identification of antioxidant and α-glucosidase inhibitory compounds from fruit juice of Nitraria tangutorum. Food Chemistry 227:93−101

doi: 10.1016/j.foodchem.2017.01.031
[66]

Bakri M, Yi Y, Chen LD, Aisa HA, Wang MH. 2014. Alkaloids of Nitraria sibirica Pall. decrease hypertension and albuminuria in angiotensin II-salt hypertension. Chinese Journal of Natural Medicines 12:266−272

doi: 10.1016/s1875-5364(14)60053-4
[67]

Turghun C, Bakri M, Abudulla R, Sun G, Aisa HA. 2018. UHPLC-MSn -assisted characterization of bioactive alkaloids extracted from Nitraria sibirica leaves and enriched using response surface method and adsorption on macroporous resin. Industrial Crops and Products 125:529−536

doi: 10.1016/j.indcrop.2018.09.038
[68]

Zhao B, Liu J, Chen X, Zhang J, Wang J. 2018. Purification, structure and anti-oxidation of polysaccharides from the fruit of Nitraria tangutorum Bobr. RSC Advances 8:11731−11743

doi: 10.1039/c8ra01125g
[69]

Abuduwaili A, Zhang X, Yi Y, Yang B, Wali A, et al. 2026. A comparative study on the structural properties of polysaccharides from Nitraria sibirica fruits with different extraction methods and their antioxidant and prebiotic activities. Chemistry & Biodiversity 23:e02596

doi: 10.1002/cbdv.202502596
[70]

Boubaker J, Skandrani I, Bouhlel I, Ben sghaier M, Neffati A, et al. 2010. Mutagenic, antimutagenic and antioxidant potency of leaf extracts from Nitraria retusa. Food and Chemical Toxicology 48:2283−2290

doi: 10.1016/j.fct.2010.05.061
[71]

Ma T, Hu N, Ding C, Zhang Q, Li W, et al. 2016. In vitro and in vivo biological activities of anthocyanins from Nitraria tangutorun Bobr. fruits. Food Chemistry 194:296−303

doi: 10.1016/j.foodchem.2015.07.110
[72]

Chaâbane M, Koubaa M, Soudani N, Elwej A, Grati M, et al. 2017. Nitraria retusa fruit prevents penconazole-induced kidney injury in adult rats through modulation of oxidative stress and histopathological changes. Pharmaceutical Biology 55:1061−1073

doi: 10.1080/13880209.2016.1278455
[73]

Du X, Cheng X, Feng M, Liang Y, Wang Y, et al. 2025. Nitraria tangutorum fruit extract modulates rumen fermentation and microbiota in Hu sheep. Journal of Agriculture and Food Research 24:102282

doi: 10.1016/j.jafr.2025.102282
[74]

Boubaker J, Bhouri W, Ben Sghaier M, Bouhlel I, Skandrani I, et al. 2011. Leaf extracts from Nitraria retusa promote cell population growth of human cancer cells by inducing apoptosis. Cancer Cell International 11:37−45

doi: 10.1186/1475-2867-11-37
[75]

Boubaker J, Lahmar A, Ben Toumia I, Ghedira K, Chekir-Ghedira L. 2024. Antitumoral potency of isorhamnetin and its derivates carbohydrate in methanol extract from Nitraria retusa leaves. Research Square Preprint

doi: 10.21203/rs.3.rs-5395825/v1
[76]

Boubaker J, Wissem B, Mohammed BS, Ines B, Mounira K, et al. 2012. Flavonoids products from Nitraria retusa leaves promote lymphoblastoid cells apoptosis. Nutrition and Cancer 64:1095−1102

doi: 10.1080/01635581.2012.717680
[77]

Wang Y, Xie Q, Ding X, Sun X, Luo S, et al. 2018. 白刺对胃癌MGC-803细胞的抑制作用[Study of Inhibitory Action of BaiCi on Gastric Cancer MGC-803 Cells]. 西部中医药 [Western Journal of Traditional Chinese Medicine] 31:14−18 (in Chinese)

[78]

Liu BPL, Chong EYY, Cheung FWK, Duan JA, Che CT, et al. 2005. Tangutorine induces p21 expression and abnormal mitosis in human colon cancer HT-29 cells. Biochemical Pharmacology 70:287−299

doi: 10.1016/j.bcp.2005.04.024
[79]

Yu F, Wei K, Lian X. 2003. Effect and mechanism of nitrary flavone on Hep, U_ (14) tumor of mice. Chinese Pharmacological Bulletin 19:886−888

[80]

Boubaker J, Chaabane F, Bedoui A, Aloui R, Ben Ahmed B, et al. 2015. Antitumoral potency of methanolic extract from Nitraria retusa leaves via its immunomodulatory effect. Cancer Cell International 15:82

doi: 10.1186/s12935-015-0232-y
[81]

Boubaker J, Ben Toumia I, Sassi A, Bzouich-Mokded I, Ghoul Mazgar S, et al. 2018. Antitumoral potency by immunomodulation of chloroform extract from leaves of Nitraria retusa, Tunisian medicinal plant, via its major compounds β-sitosterol and palmitic acid in BALB/c mice bearing induced tumor. Nutrition and Cancer 70:650−662

doi: 10.1080/01635581.2018.1460683
[82]

Zar Kalai F, Han J, Ksouri R, El Omri A, Abdelly C, et al. 2013. Antiobesity effects of an edible halophyte Nitraria retusa forssk in 3T3-L1 preadipocyte differentiation and in C57B6J/L mice fed a high fat diet-induced obesity. Evidence-Based Complementary and Alternative Medicine 2013:368658

doi: 10.1155/2013/368658
[83]

Li B, Liu K, Zhang J, Li C. 2019. Anthocyanins from the fruits of Nitraria tangutorun Bobr. improve nonalcoholic fatty liver injury induced by high-fat diet in mice. Food Science 40:217−223

doi: 10.7506/spkx1002-6630-20180913-138
[84]

Zar Kalai F, Han J, Ksouri R, Abdelly C, Isoda H. 2014. Oral administration of Nitraria retusa ethanolic extract enhances hepatic lipid metabolism in db/db mice model 'BKS. Cg-Dock7m+/+ Leprdb/J' through the modulation of lipogenesis-lipolysis balance. Food and Chemical Toxicology 72:247−256

doi: 10.1016/j.fct.2014.07.029
[85]

Du X, Cheng X, Dong Q, Zhou J, Degen AA, et al. 2022. Dietary supplementation of fruit from Nitraria tangutorum improved immunity and abundance of beneficial ruminal bacteria in Hu sheep. Animals 12:3211

doi: 10.3390/ani12223211
[86]

Iida A, Usui T, Zar Kalai F, Han J, Isoda H, et al. 2015. Protective effects of Nitraria retusa extract and its constituent isorhamnetin against amyloid β-induced cytotoxicity and amyloid β aggregation. Bioscience, Biotechnology, and Biochemistry 79:1548−1551

doi: 10.1080/09168451.2015.1027655
[87]

Wang H, Zhou J, Bi H, Yang X, Chen W, et al. 2021. Bioactive ingredients from Nitraria tangutorun Bobr. protect against cerebral ischemia/reperfusion injury through attenuation of oxidative stress and the inflammatory response. Journal of Medicinal Food 24:686−696

doi: 10.1089/jmf.2020.4848
[88]

Kurskaya O, Prokopyeva E, Bi H, Sobolev I, Murashkina T, et al. 2022. Anti-influenza activity of medicinal material extracts from Qinghai–Tibet plateau. Viruses 14:360

doi: 10.3390/v14020360
[89]

Mei X, Dai T, Shen Y. 2023. Adaptive strategy of Nitraria sibirica to transient salt, alkali and osmotic stresses via the alteration of Na+/K+ fluxes around root tips. Journal of Forestry Research 34:425−432

doi: 10.1007/s11676-022-01486-1
[90]

Zhu L, Li M, Huo J, Lian Z, Liu Y, et al. 2021. Overexpression of NtSOS2 from halophyte plant N. tangutorum enhances tolerance to salt stress in Arabidopsis. Frontiers in Plant Science 12:716855

doi: 10.3389/fpls.2021.716855
[91]

Duan R, Zhang H, Zhao Y, Zhang H, Li R, et al. 2025. Analysis of SRO gene family in Nitraria sibirica Pall. and the function of NsSRO1a in improving plant drought tolerance. Tree Physiology 45:tpaf050

doi: 10.1093/treephys/tpaf050
[92]

Lu L, Chen X, Wang P, Lu Y, Zhang J, et al. 2021. CIPK11: a calcineurin B-like protein-interacting protein kinase from Nitraria tangutorum, confers tolerance to salt and drought in Arabidopsis. BMC Plant Biology 21:123

doi: 10.1186/s12870-021-02878-x
[93]

Zhang H, Qi C, Li C, Huang D, Mao H, et al. 2024. Overexpression of high affinity K+ transporter from Nitraria sibirica enhanced salt tolerance of transgenic plants. Plant Science 342:112052

doi: 10.1016/j.plantsci.2024.112052
[94]

Qi D, Xiao H, Liu H, Zheng L, Wang Y. 2025. Sucrose enhances adventitious root formation in Nitraria tangutorum under drought stress via hormonal homeostasis and hydrogen peroxide signaling. Physiologia Plantarum 177:e70521

doi: 10.1111/ppl.70521
[95]

Zhang H, Hu A, Wu H, Zhu J, Zhang J, et al. 2023. Integrated metabolome and transcriptome analysis unveils novel pathway involved in the fruit coloration of Nitraria tangutorum Bobr. BMC Plant Biology 23:65

doi: 10.1186/s12870-023-04076-3
[96]

Bao X, Zong Y, Hu N, Liu B, Wang H. 2023. 西伯利亚白刺NsMYB5调控果实花青素生物合成[NsMYB5 regulates anthocyanin biosynthesis in fruits of Nitraria sibirica]. 西北农业学报 [Acta Agriculturae Boreali-occidentalis Sinica] 8:1215−1222 (in Chinese)

doi: 10.7606/j.issn.1004-1389.2023.08.008
[97]

Gravel E, Harfouche A, Salame R, Leblanc K, Maciuk A, et al. 2013. Spontaneous formation of nitrarine and polycyclic skeletons related to Nitraria indolic alkaloids under non-enzymic conditions. Chemistry – A European Journal 19:14515−14520

doi: 10.1002/chem.201301877
[98]

Granot G, Grafi G. 2014. Epigenetic information can reveal phylogenetic relationships within Zygophyllales. Plant Systematics and Evolution 300:1819−1824

doi: 10.1007/s00606-014-1008-x