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2022 Volume 2
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Native and xeric plant recommendations for urban landscapes in Kuwait

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  • The utilization of native and xeric plants in urban landscapes has gained great interest in recent years due to two pressing issues; climate change effects and water scarcity. These two issues are further exacerbated by Kuwait’s arid desert environment, poor soil and salinity, making it quite challenging for popular ornamental plants to flourish in urban landscapes. Kuwait is currently dominated by a European aesthetic in terms of urban green spaces, which include plants such as Concarpus erectus, Nerium oleander, Vitex purpurea, Vitex trifolia and Bougainvillea glabra, all of which are not compatible with Kuwait's harsh climate and are environmentally unsustainable. This paper discusses climatic conditions, soil characteristics and water resources that are available and includes a plant list that has been created of recommended native and non-native xeric species that are capable of withstanding Kuwait's harsh climatic conditions to create aesthetically pleasing and sustainable green urban landscapes.
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  • Cite this article

    Almutawa AA. 2022. Native and xeric plant recommendations for urban landscapes in Kuwait. Technology in Horticulture 2:7 doi: 10.48130/TIH-2022-0007
    Almutawa AA. 2022. Native and xeric plant recommendations for urban landscapes in Kuwait. Technology in Horticulture 2:7 doi: 10.48130/TIH-2022-0007

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Native and xeric plant recommendations for urban landscapes in Kuwait

Technology in Horticulture  2 Article number: 7  (2022)  |  Cite this article

Abstract: The utilization of native and xeric plants in urban landscapes has gained great interest in recent years due to two pressing issues; climate change effects and water scarcity. These two issues are further exacerbated by Kuwait’s arid desert environment, poor soil and salinity, making it quite challenging for popular ornamental plants to flourish in urban landscapes. Kuwait is currently dominated by a European aesthetic in terms of urban green spaces, which include plants such as Concarpus erectus, Nerium oleander, Vitex purpurea, Vitex trifolia and Bougainvillea glabra, all of which are not compatible with Kuwait's harsh climate and are environmentally unsustainable. This paper discusses climatic conditions, soil characteristics and water resources that are available and includes a plant list that has been created of recommended native and non-native xeric species that are capable of withstanding Kuwait's harsh climatic conditions to create aesthetically pleasing and sustainable green urban landscapes.

    • Growing ornamental plants in Kuwait is already quite challenging due to its harsh environmental conditions[1]. In addition, the effects of global climate change are continuously accelerating, therefore effective biological responses need to be in place to guide the effective protection of biodiversity[2]. Already less than 10% of Kuwait is covered with natural vegetation and there is evidence that the geographical distribution of dominant perennial plants in Kuwait is being impacted by climate change[1,36]. Moreover, further natural landscapes are being lost as a consequence of altered land use as a result of population growth that has led to an increase in urbanization and industrialization. Altered land use is one of the main drivers of biodiversity depletion and habitat loss, pushing native plant populations to regional extinction[7]. Additional activities that contribute to native plant loss are overgrazing, recreational activities (off-road driving and camping), industrial practices (specifically quarrying) and the remnant effects of the Gulf War[79]. Efforts have been dedicated toward the rehabilitation and conservation of Kuwait's desert ecosystem. Of the total territorial area in Kuwait approximately, 11% is either terrestrial or marine protected areas[10]. Several protected terrestrial areas that have been established in Kuwait include Sabah Al-Ahmad natural reserve, Jahra nature reserve, Doha reserve, Kuwait Institute of Scientific Research (KISR) Sulaybia Field Research Station, Al-Kabid nature reserve, Al-Khuwaisat, Al-Sulaibikhat nature reserve, Om-igdar, Khobar Al-awazim, Wadi Al-Babtain, Qurain Hill nature reserve, Al-Abdaliya nature and Al-Leyah protected area[11]. However, the majority of these protected areas are situated away from urban landscapes and are located in the desert. Only one project incorporated native species on a large scale in Kuwait, and that is at Bawabat Al-Shamiyah. Therefore, there is a need to connect these native habitats to urban areas to combat urban habitat fragmentation.

      Native plants are defined as plants that are a part of the balance of nature that has developed over hundreds or thousands of years in a particular region or ecosystem[12]. The utilization of native species in their natural habitat and xeric plants acclimatized to the region's geo-climatic conditions have been encouraged. As they have a higher survival rate, adaptation potential, lower maintenance, disease resistance, salinity tolerance, nitrogen fixation ability, ethnobotanical importance and medicinal uses[13]. These plants can help create a sustainable landscape that can address habitat fragmentation and habitat loss through xeric landscaping. Xeric landscaping refers to low water use landscaping and water-efficient landscaping that potentially offers a balance of water-efficient landscapes without soil water limits[14,15]. This method of landscaping has been widely applied in Australia, South Africa and Arizona, USA, where alternative man-made 'wild' nature is used in opposition to the more conventional 'obedient' landscapes[16]. Xeric plant species can be native and non-native species that have drought survival adaptive mechanisms[14,17]. These adaptive mechanisms encompass extensive root systems, thick cuticles, stomatal openings that are protected and the occurrence of bulliform cells[14,17,18]. Low transpiration rates through small leaves lead to less evaporation of water due to the reduced total surface area of the leaf. Thick waxy cuticles, leathery or glossy cuticles that act as a barrier to evaporation and reflect sunlight[14,17]. Their main growing period is winter and spring when rain is abundant[19]. Bulliform cells are bubble-shaped epidermal cells that cause leaves to roll or fold in harsh environmental conditions by losing their turgor pressure in water stress conditions preventing moisture loss through the vacuoles and reopening again under favourable conditions[18,20]. Bulliform cells are especially associated with Poaceae, Cyperaceae and Juncaceae species[18,20].

      The plant selection process discussed in this paper covers a range of topics from the climatic conditions, and soil and water conditions in Kuwait, native and non-native species and annual and perennial species. All the plants discussed are native plants or introduced plants that have been studied, suggested or utilized by KISR and the Public Authority of Agriculture and Fisheries, Kuwait (PAAF) to give a more comprehensive overview of tackling urban landscaping that is distinct to the region.

    • As a desert environment, Kuwait experiences harsh climatic conditions, with prolonged periods of extreme heat (Table 1). During the summer months, the average temperature is 46.22 °C, while in the winter months, it can drop to 6 °C[21,22]. Evaporation rates in summer range from 3.1 to 24.3 mm/day, despite low humidity (13%)[2325]. Precipitation is minimal, occurring in the spring and increasing in frequency around November, averaging 22 wet days which amounts to 115 mm, annually fluctuating between 25 and 250 mm[24,26]. The wind blows predominantly during the summer from the northwest, forming 60% of the total wind direction and from the southeast to a lesser extent and shorter duration[26]. Therefore, the dust activity in Kuwait, is low during winter, growing strong between March and April, and at its highest in June and July[27]. The average monthly wind speed in June reaches a maximum of 20.8 km h−1 and a minimum of 12.1 km h−1 in November[9]. Kuwait experiences an average of 3347 sunshine hr/year. The number of sunshine hours ranges; from an average of 7 hr/day in December, and 11 hr/day in August[28].

      Table 1.  Monthly climate and weather averages in Kuwait (adapted from www.timeanddate.com/weather/kuwait/kuwait-city/climate).

      Month
      JanFebMarAprMayJunJulAugSepOctNovDec
      High temperature (°C)192227334044464642362721
      Low temperature (°C)791319252930302621149
      Mean temperature (°C)131520263236383834282015
      Percipitation (mm)10.83.72.315.711.810.118.43.71.70.30.94.1
      Humidity64%54%43%34%21%13%14%18%21%36%50%62%
      Wind (km/h)121414151520191614121212

      It is inevitable that any long-term changes in climate patterns will have an impact on vegetation type and canopy structure, as well as bare land, which will have an impact on land-surface albedo and evapotranspiration, modifying near-ground temperatures and precipitation[3,29,30]. Future changes in vegetation cover could have a similar impact on rainfall as climates change, but the magnitude of those changes is uncertain[3,31].

    • Kuwait's topography consists of predominantly flat and gently undulating land that sparsely encounters elevations, wadies (drainage lines), depressions, sand dunes and salt marshes[32,33]. The land rises gradually from the shores of the Arabian Gulf from an altitude of zero to approximately 300 m southwest of the country[32]. There are eight soil groups that dominate Kuwait[34]: spetrogypsids, torripsamments, haplocalcids, haplogypsids, aquisalids, calcigypsids, petrogypsids and torriorthents (Fig. 1, Table 2)[34, 35].

      Figure 1. 

      Map indicating Kuwait soil group classification spread (data from KISR[37]; PAAFR, 1999).

      Table 2.  Kuwait soil group classification (adapted from Shahid & Omar[35]; Omar et al.[34]; Soil Survey Staff [36]; KISR[37]; PAAFR, 1999).

      Soil groupsDescription
      TorripsammentsWell to somewhat excessively drained, deep or very deep sandy soils.
      PetrocalcidsWell drained or moderately drained, shallow or moderately deep, sandy to loamy soils overlying a calcic hardpan. When upper soil is truncated, it may appear at the surface.
      HaplocalcidsWell drained, deep or very deep, sandy to loamy soils, which have a layer ofcarbonate masses and nodules in the profile.
      *HaplogypsidsWell drained, deep or very deep, sandy to loamy soils, which have a layer of gypsum crystals in the profile.
      AquisalidsPoorly or somewhat poorly drained, deep or very deep, sandy to clayey soils. Within the soil there is a layer of salt accumulation that usually occurs near the surface.
      CalcigypsidsWell drained, deep or very deep, sandy to loamy soils containing a layer of carbonate masses and nodules and a layer of gypsum crystals within the profile.
      PetrogypsidsWell drained, shallow or moderately deep, sandy to loamy soils overlying a gypsic hardpan. Hardpan may be exposed at surface, when upper soil is truncated.
      TorriorthentsExcessively drained to well drained, moderately deep or very deep, sandy soils. Within the soil profile there is a high content of shell fragments and some gypsum accumulations
      MiscellanousRefers to areas that are incapable of supporting vegetation without significant reclamation efforts. This includes dumps and quarries and urban and industrial areas.
      *Due to map scale limitations, Haplogypsids are not shown.

      In many parts of Kuwait, soil is partially consolidated with a sediment of a massive calcrete type, known locally as 'gatch'[3840]. It is a caliche layer (duripan or hardpan) that is found in varying depths but on average at a 2 m depth, attaining a thickness of tens of meters[22,38,40]. Gatch consists of a mixture of quartz sands, silica and gypsum, cemented predominantly by carbonates (calcite and/or dolomite)[22,38,40]. This gatch zone decreases in porosity and effective permeability due to cementation by carbonate[22]. Gatch soils are considered impervious and require proper drainage arrangements when used as fill material as the average shrinkage limit is 17% above normal moisture content. Therefore, soils are liable to swell when saturated and crack when dry[22]. However, soil stability can be through the addition of natural and synthetic chemicals that can improve the physical and chemical soil properties through aggregate stability by enhancing soil strength, aggregate stability, and water infiltration. They also reduce the effects of salinity on plant growth and development and supply nutrients[41, 43]. Additionally, a study on the development and evaluation of a new multi-purpose soil additive by Lahaliah[39], determined the addition of 1.0% sulfonated urea-melamine-formaldehyde (SUMF) improved the compressive strength of gatch sand from 4.74 to 13.38 kg cm−2[42]. The use of a mixture of a polyanionic polymer (sulfonated amino formaldehyde, polyanionic condensates) with polyvinyl alcohol was established to achieve very good stability for gatch, however, it is expensive and difficult to dissolve in water[42]. Another more economical and time-feasible option is to drill holes at selected sites where trees or deep-rooted species will be planted. Growing vegetation on gatch soil has been successful in the past, as proven by Suleiman et al.[40] in a study on plant enrichment in desert ecosystems in Kuwait, with relative growths rates of 113.2% for Pennisetum divisum, 67.8% for Calligonum polygonoides, 51.5% for Farsetia aegyptia, 30.1% for Helianthemum lippii, 12.8% for Tephrosia haussknechtii, 7.9% for Prosopis juliflora and 6.9% for Prosopis spicigera, with the exception of Tephrosia haussknechtii and Rhanterium epapposum, which had survival rates of less than 80%[43].

      It must be noted that, due to urbanization, landscape soils are usually physically and chemically altered, and the microclimate in those spaces has been transformed[43]. Moderate to severe land degradation and plant cover deteriorated, affecting around 75% of Kuwait's desert as a result of the high intensity of anthropogenic activities[7,9,44,45]. Soil can be improved to promote plant survival and germination rate through the addition of mulch and fertilizer for selected plants. Nevertheless, since there is not a universal solution, therefore plant palette selection for the selected site needs to be determined ahead of soil preparation.

    • Kuwait has limited natural water resources, groundwater is the only natural water resource it has, with the exception of isolated freshwater lenses in Raudhatain and Umm Al-Aish in Northern Kuwait[46]. The rest of the available groundwater is of brackish quality, with the majority of it located in the western and central regions of Kuwait. The brackish water from these locations is utilized for landscaping and agricultural purposes, buildings and road construction, and is blended with desalinated seawater[21,46]. Brackish water has a salt concentration that ranges from 3,000 to 8,000 mg L−1, which limits plant selection[21]. The irrigation of landscape plants using treated municipal sewage water and industrial water, subjecting it to tertiary treatment has become a common practice[21,25]. The salt concentration is 1,300−2,430 mgL−1 and contains both organic and inorganic matter[25].

      Irrigation is a crucial part of many landscapes and important to maintaining healthy plants. To combat the dry climate in Kuwait and the low water holding capacity of soil, excessive irrigation is a common practice in landscaping[47]. To ensure efficient water use, drip lines should be used instead of furrows and sprinklers as a method of irrigation. During summer, drip irrigation is recommended twice weekly; during winter, drip irrigation is recommended daily depending on the plant's water requirements[43].

    • In the 1960s Kuwait developed the first National Greening Plan (NGP)[21,48]. The aims of the NGP were: the beautification of Kuwait City and its suburbs, the rehabilitation of deteriorating rangelands, the establishment of green parks in the desert, the expansion of existing agricultural areas, the creation of plantations or artificial forests, and enriching the coastline with mangroves[48,49]. Exotic species, such as Eucalyptus, Tamarix, Ziziphus trees and Jasminum and Lantana shrubs, were utilized during that time in urban spaces[21]. Currently, green urban landscapes are made up predominantly of exotic species from temperate and semi-temperate regions: examples include Concarpus erectus, Nerium oleander, Vitex purpurea, Vitex trifolia and Bougainvillea. However, greening these urban spaces using the current limited regional and commercially available plant palette is generating a variety of ecological problems. These ecological problems include the risk of invasive plants, dominant plant populations taking over, changing the local ecosystem and having high-water requirements[50,51].

    • Residential landscapes that reflect local environmental conditions by utilizing native plant species for ornamental purposes can be a more viable option, as they offer a variety of benefits that maximize the future well-being of people and the natural environment[52]. Native plant species are well adapted to the regional climatic conditions, have the capacity to adapt to adverse local environmental conditions, and have low water and maintenance requirements[43,50,52]. They also promote the biodiversity of local native wildlife in Kuwait by preserving the ecological balance between native plants and fauna[43,52]. Specific native plants have been selected in order to avoid any landscape design problems that may arise, including seasonal aesthetics, mass, form and color palette (Table 3). However, if locally adapted species become isolated and cannot colonize new habitats, they may not be able to shift their ranges with climate change[2]. Additionally, native plants are not allowed to be commercially sold, propagated or collected by nurseries in Kuwait due to legislation stated in the list of unauthorized traded seeds and tubers created by PAAF.

      Table 3.  Recommended native plant species for green urban landscapes in Kuwait.

      Vegetation formPlant nameFamilyDescriptionPlant spacingFlowering timeEnvironmental conditionsAdditional info
      ShrubsHelianthemum lippi Rakrouk
      (Fig. 2a)
      CistaceaeSmall perennial shrubby plant 10−45 cm tall[60]Triangular spacing
      of 2 m[55]
      March−AprilOccurs on sandy soils and sandy limestone gypsum[60]• Used in the stabilization of sand.
      • Host plant for several species of desert truffles.
      • Medicinal properties[60].
      Ziziphus nummularia
      Jujube
      (Fig. 2b)
      Rhamnaceae
      Thorny shrub that grows to 1−5 m height. Branches zig-zag, covered with white coating. Stems and branches are purplish to ashy. Bark light brown. Multi-stemmed from the base leaves shortly stalked, ovate to orbicular, serrate, 12−18 mm long, 8−10 mm broad, round at apex and base, more or less tomentose above but densely wooly beneath. Stipular spines in pairs, one slender and straight whereas other short and recurved. Flowers cream color, flowers during July- August and the ripe fruits are available in November-December[56]6 m × 6 m[56]March−June
      Hardy enough to grow not only in hostile arid environment but also in poor and different forms of soil profiles underneath[56]. It is adapted to flat alluvial plains with fairly deep (80-120 cm) sandy loam soils and with a hard kankar pan below[56]. Flat alluvial plains with sandy clay loam to clayey soils and a kankar pan at 45-60 cm below the soil also show high shrub density in semi-arid regions[56]. Drought hardy, thrives in areas with 125 to 850 mm rainfall[56].• Fodder yielding shrub.
      • Source of fruit.
      • Source of wood.
      • Medicinal uses: bark is used in diarrhea, while roots are used to make digestive and nutritive tonic[56].
      Farsetia aegyptia

      Al-labana
      (Fig. 2c)
      Brassicaceae
      Native grey-green woody perennial about 30 cm in height. It has slender, smooth and multi-branched stems. The flowers are creamy brown with four petals. Two rows of seeds are formed in an oval- shaped seed pod. It flowers in April[61]Triangular
      spacing of 2 m[55]
      March−AprilGrows on gravelly soils to stony ground. It grows in severe climatic conditions with under 100 mm rainfall per year[66].
      • Traditionally used to treat toothache and gingivitis, and for sore eyes.
      • A decoction of the plant mixed with alum is used as a mouth rinse and for disinfecting (Morocco)[66].
      Haloxylon
      salicornicum Rimth
      (Fig. 2d)
      Amaranthaceae
      Dwarf branching perennial shrub, with a plant height of 0.75–1.0 m. Bark is pale in color, stem and branches are pale yellow, jointed producing two triangular scales which take the place of leaves. Branches are erect or ascending terete. Almost leafless and leaves reduced to dilated tips of the joints. Has a wide-spread and deep tap root system that extends beyond 5 m and estimated to go 8–10 m deep. Flowers are bisexual and born singly by short twigs in axes of scale like bracts[58]2 mSept−OctIs resilient to a variety of habitats: shallow runnels, alluvial fans, shallow silt, low dunes, hard gravel and sandy gravelly wadi-terraces, sandy plains, gravel desert, sandy undulating plains, sandy plains and inter-dunal areas. It requires a minimum annual rainfall of 100 mm[5]. It can tolerate high salinity up to 8,000–10,000 mg L−1[58].• Fruiting tops and seed as animal feed.
      • Seeds as emergency food.
      • Wood as fuel.
      • Different plant parts for medicine.
      • Used in restoration of degraded lands[58]
      Lycium shawii
      Roem. & Schult.
      Awsaj
      (Fig. 2e)
      Solanaceae
      Perennial shrub with purple, sometimes white, trumpet like flowers and sharp thorns. Leaves are elliptical and congested in closed clusters. Fruits are globular, red to orange berries which are edible [59]2 m × 2 m[62]March−April
      and
      throughout
      the year in
      irrigated soil
      Grows in a variety of desert habitats; gravel plains, hillsides, valleys, sandy stone ridges and along drainage basin, such as valleys and depressions[67].• Fruits are edible.
      • provides honey for wild bees.
      • Food and shelter for wild birds and animals.
      • The roots are boiled and the decoction is used to treat sores in the mouth, coughs, backache and administered internally to cure tick fever in livestock.
      • Leaves are used to treat constipation and stomach ache.
      • Leaves are much liked by livestock[59].
      Ochradenus baccatus Delile

      Taily Weed
      (Fig. 2f)
      Resedaceae
      Perennial shrub, grows up to 2 m high, with grey green linear leaves, blooms yellow flowers, followed by white berries. Stems greenish in color[57].1.5 m × 1.5 m[62]
      Dec−MarchGrows on dry soils and colonizes arid environments where saline soils are common. Has high drought tolerance[68]• Fruits are one of the most important food sources for many animal species in the Arabian desert.
      • O. baccatus have various medicinal uses.
      • Useful in controlling root-knot nematodes.
      • Planted as cover plant/ green manure[68].
      Calligonum polygonoides
      Erta
      (Fig. 3a)
      PolygonaceaeSmall perennial shrub that is about 1 to 2 m high, with a girth of 30−60 cm. Stem has nodes and internodes which join stem and branches conspicuously. Flowers: are white, small, bisexual and regular. Fruit is oblong and nut-like; small and rounded seeds[63].Triangular spacing
      of 2 m[55]
      March−AprilGrows on dry sandy soil of the desert[69]. It is highly resistant to drought [70].
      • Harvested for use as fodder.
      • Fuel wood.
      • Food.
      • Used for sand dune stabilization[70]
      Heliotropium
      bacciferum
      Salt heliotrope
      (Fig. 3b)
      BoraginaceaeAscending or procumbent, herbaceous perennial undershrub, that grows up to 50 cm. Dark green with semi-fleshy leaves and white bristles. Flowers, small, white, tubular; fruits achenes[74].30–60 cm[75]March−AprilGrows in saline sandy soils, sandy wadis, on calcareous ridges. Dominate disturbed locations and roadsides[74].• Dried and powdered plant is added to water and drunk to combat fatigue.
      • Leaf sap is applied to burns.
      • Applied topically to treat headache.
      • Used internally to treat gonorrhea and to increase lactation.
      • Macerate of the plant is used as ink[74].
      GrassesPennisetum divisum
      Thammam
      (Fig. 3c)
      PoaceaePerennial grass. It can grow quite large in substantial clumps up to 1−3 m in height[64].Triangular spacing
      of 2 m[55]. Other options include;
      90 m × 1.20 m, 50 cm × 50 cm, depending on aesthetic.
      Feb−AprilDrought-resistant, grows in depressions and shallow wadis with coarser textured sandy and gravelly soil[71]. Naturally found on torripsamment and aquisalid soil types, located on level coastal plains covered by uneven sand sheets or isolated hummocks[34].• Animal Fodder.
      • Used in land stabilization for soil protection and sand dune fixation[71].
      Panicum turgidum
      Taman, tuman, or thaman
      (Fig. 3d)
      Poaceae Perennial growing to 1.5 m × 0.2 m at a fast rate[65].Triangular spacing
      of 2 m[55]. Other option include; 90 cm × 120 cm or 50 cm × 50 cm, depending on aesthetic.
      Feb−AprilDrought-resistant, grows in shallow wadis and runnels in deep sandy soil, salt-affected areas and is a salt excluder[71, 72]. Naturally found on torripsamment and aquisalid soil types, located on level coastal plains covered by uneven sand sheets or isolated hummocks[34].• Animal Fodder.
      • Used in land stabilization for soil protection and sand dune fixation[71].
      Ground
      cover
      Cornulaca aucheri
      Hadh, saley, or thalj
      (Fig. 3e)
      Amarantha
      ceae
      Annual or short-lived perennial, leaves are linear-filiform to triangular-subulate, long, greyish-green, re-curved, white spine and densely woolly in axils. Average length is 94.2 cm in average length, average height is 141.7 cm[73].-April−MayGrows in wadi, depressions, and playas[73]. Have a short life span and dominate immediately after a disturbance[34].• Used for sand fixation and stabilization[73].
      • Used as camel fodder[119].
      Salsola imbricata
      Saltwort, milayah
      (Fig. 3f)
      Amarantha
      ceae
      Succulent light green undershrub with rudimentary leaves on red and green branches. Flowers minute, green, bisexual with 5 stamens; calyces inflated in fruit forming transparent wings surrounding the fruitJune−SeptDisturbed areas, saline soils, sandy desert and coastal areas[119]. Have a short life span and dominate immediately after a disturbance[34]. Naturally found on aquisalids soil types[34].• Traditionally being used to treat asthma, cough and congestion.

      Figure 2. 

      (a) Helianthemum lippi (Rakrouk), (b) Ziziphus nummularia (Jujube), (c) Farsetia aegyptia (Allabana), (d) Haloxylon salicornicum (Rimth), (e) Lycium shawii Roem. & Schult. (Awsaj), (f) Ochradenus baccatus Delile (Taily weed).

      Figure 3. 

      (a) Calligonum polygonoides (Erta), (b) Heliotropium bacciferum (Salt heliotrope), (c) Pennisetum divisum (Thammam), (d) Panicum turgidum (Taman, tuman, or thaman), (e) Cornulaca aucheri (Hadh, saley, or thalj), (f) Salsola imbricata (Saltwort, milayah).

      Green urban landscapes do not have to encompass strictly native plant species to conserve resources. Incorporating non-native or naturalized species that are adapted to Kuwait's climatic conditions can provide economic landscape potential and a broader range of color palettes and aesthetics[53]. Drought-tolerant species have specific characteristics that enable them to withstand extreme drought and desert conditions. These species can shed their leaves to further decrease water loss in severe drought conditions[54]. In order to provide recommendations for future landscape projects in Kuwait, a list of previous research studies and projects in countries with similar climatic conditions and within the same region have been compiled (Table 4).

      Table 4.  Recommended non-native xeric plant species that are tolerant of Kuwait’s climatic conditions.

      Vegetation formPlant nameFamilyNative toDescriptionExposureEnvironmental conditionsAdditional info
      Trees Eucalyptus camaldulensis DEHNH
      River red gum
      MyrtaceaeAustraliaEvergreen single-stemmed, large-boled, medium-sized tree, commonly grows to about 20 m tall, though occasional specimens can reach
      50 m[80].
      Full SunHigh salinity tolerance, medium drought tolerance, medium irrigation is required[19]. It grows in areas where the mean annual precipitation is in the range of 250−2,500 mm and an annual temp. range of 3−22 °C min to 21−40 °C max[80]. Plant prefers a pH in the range 5−7, tolerating 4.5−8[80]. Reported to grow in areas with only 200 mm rainfall, Succeeds in most soils[80].• Leaves are used as an antiseptic and is used for relieving coughs and colds, sore throats and other infections.
      • Plant is an aromatic, astringent, tonic herb that sticks to the teeth and turns the saliva red.
      • Resin contains tannins and is used internally in the treatment of diarrhea and bladder inflammation, externally it is applied to cuts.
      • Smoke from burning leaves helps to repel insects[80].
      Eucalyptus microtheca F. MUELL Coolibah
      MyrtaceaeAustraliaEvergreen shrub or small tree grows to 3−20 m tall depending on growing conditions[79].Full sunHigh salinity tolerance, medium drought tolerance, low irrigation is required[19]. Tolerant to mean annual rainfall in the range 250−1,000 mm, but tolerates 150−1,200 mm[79]. A pH in the range 6.5−8, tolerating 6−9[79].• Planted for shelterbelts, shade, soil conservation, and erosion control.
      • Flowers are an excellent source of nectar and pollen for bees.
      • Inner bark is a traditional Aboriginal medicine, mashed and prepared into a poultice to treat snakebites.
      • Seeds are powdered and made into cakes.
      • Source of water where no free water was available[79].
      Acacia rigens

      Needle bush wattle
      Fabaceae
      AustraliaCompact, rounded shrub grows from 2.4 to 3 m tall, bright yellow flowers[76].Full sunDoes well with dry conditions and a well-drained soil[76].• Provide: fuel, firewood, timber, forage, gum, tannins, fiber, folk medicine, and food.
      • useful for environmental protection and soil and water conservation.
      • Support large numbers of herbivorous vertebrates and invertebrates and nectarivorous insects[82].
      Acacia saligna


      Golden wreath wattle
      Fabaceae
      AustraliaSmall tree grows from 3 to 7.6 m tall with a bushy, spreading crown, Golden yellow to almost orange flowers[76].Full sunIt does well in almost any soil, from light to heavy[76].• Provide: fuel, firewood, timber, forage, gum, tannins, fiber, folk medicine, and food.
      • useful for environmental protection and soil and water conservation.
      • Support large numbers of herbivorous vertebrates and invertebrates and nectarivorous insects[82].
      Acacia ehrenbergiana Hayne.

      Tamat

      Fabaceae
      Mauritania, Senegal, Mali, Upper Volta and Niger, and extending into the Sahara and to the Nile Valley.
      4.5 m high
      Full sunDrought tolerant species, can survive in areas that experience rainfall of 50−400 mm/annum[82].• Provide: fuel, firewood, timber, forage, gum, tannins, fiber, folk medicine, and food.
      • useful for environmental protection and soil and water conservation.
      • Support large numbers of herbivorous vertebrates and invertebrates and nectarivorous insects[82].
      Acacia tortilis (Forsk.) (Hayne)Fabaceae Angola, Botswana, Egypt, Eritrea, Ethiopia, Iran, Israel, Kenya, Mozambique, Namibia, Qatar, Saudi Arabia, Somalia, South Africa, Sudan, Swaziland, Tanzania, Uganda, United Arab Emirates, Zambia, Zimbabwe[99].Small to medium-sized evergreen tree or shrub that grows up to 21 m tall. Flat-topped or rounded, spreading crown; bark grey to black or dark brown, rough, fissured or smooth. Leaves glabrous to densely pubescent, glandular, short[99].Full sun Drought tolerant species, can survive in areas with rainfall of 50−400 mm/annum[82].• Provide: fuel, firewood, timber, forage, gum, tannins, fiber, folk medicine, and food.
      • useful for environmental protection and soil and water conservation.
      • Support large numbers of herbivorous vertebrates and invertebrates and nectarivorous insects[82].
      Acacia sowdenii

      Western myall
      Fabaceae
      AustraliaLong-lived, frost- and drought-resistant small tree that grows from 3.6 to 5.5 m tall with drooping branches. Yellow flowers appear irregularly, but are prolific when conditions are right[76].Full sunPrefers a dry, well-drained soil[76].• Provide: fuel, firewood, timber, forage, gum, tannins, fiber, folk medicine, and food.
      • useful for environmental protection and soil and water conservation.
      • Support large numbers of herbivorous vertebrates and invertebrates and nectarivorous insects[82].
      Prosopis cineraria


      Gaf
      Fabaceae
      Arabian gulfEvergreen tree, grows up to 6.5 m high. Leaves 1−3 jugate, glabrous or puberulous; greyish when dry[99].Full sunDrought-tolerant and thrives in hot, arid regions with an annual rainfall of less than 500 mm[76]. Suitable for: light (sandy), medium (loamy) and heavy (clay) soils, prefers well-drained soil and can grow in nutritionally poor soil that are acidic, neutral and basic (alkaline) soils, pH of 6.5−8.5, tolerating 5.5−9.8
      [78, 99].
      • Pods are used as vegetable in the dried and green form.
      • Bark is ground and used as flour.
      • Plant is reported to be astringent, demulcent, and pectoral[78].
      Ebenopsis ebano

      Texas ebony
      Fabaceae
      Texas/USAEvergreen perennial shrub or tree (1.2 m) with multiple trunks. A rounded, dense crown. Medium green, Alternate, pinnately compound leaf with 3 to 5 pairs, oblong to obovate. Small, musty fragrant, cream-colored flowers in dense, slender, terminal spikes to 3.8 cm long, flowers strongly attract bees[105].Full sunTolerant of heat and cold and full sun. Highly tolerant of desert soils. Very drought tolerant once established. Providing supplemental water to young specimens will increase growth rate[105].

      • Stipular spines that can draw blood.
      • Grows a dense canopy to the ground, crown raising is the principal pruning technique that needs to be continuously done.
      • Slow growth rate[105].

      Lysiloma watsonii

      Desert fern
      Fabaceae
      South west USALarge spreading shrub with hard brittle wood, grow up to 3.6−4.5 m. Has dense, feathery canopy, that is evergreen to partly deciduous foliage[120].

      Full sun-partial shade
      Drought tolerant species that requires full sun to partial shade. Survives in moist to dry soil, that is loam or sand texture. That is slightly acidic to highly alkaline soil in pH[120].• Requires corrective pruning to remove suckers and to help develop good structure.
      • Produces some leaf and seedpod litter
      • A good selection for the landscape when light shade is needed for smaller, understory plants.
      • Use around pools or ponds for a desert oasis theme, tropical effect, or background screen.
      • Nice patio tree or specimen for small spaces[121].
      • The bark is brewed into medicinal tea to relieve gas and symptoms of liver problems.
      • Heating the tea is used to raise the body temperature for women who are infertile because of ingestion.
      • Wood can be used to make posts and molds for building.
      • Provides shelter for animals during hot weather and nesting season[122].
      Havardia Mexicana


      Mexican ebony
      Fabaceae
      South west
      USA
      Evergreen tree, with a dark grey trunk and branch structure, grows to 12 m. Has yellow or white flowers that are very fragrant and also produces fruit and nectar that often attract wildlife[87].
      Full sun
      Prefers loose, well-drained soil with low water usage (monthly) and needs full sun for proper development. The Mexican ebony struggles in caliche or hardpan; it prefers deeper soils[87].
      • Wood is used for furniture building, as a material for agave liquor stills, firewood, and as beams for building.
      • Bark is brewed into a tea that can relieve upset stomachs.
      • Provides shade and shelter for animals.
      • Pods are edible and are often consumed by animals.
      • Fragrant flowers, fruit, and nectar attract many different forms of wildlife[87].
      Parkinsonia praecox

      Sonoran Palo Verde
      Fabaceae
      Sonora, southern Baja California into Michoacan, Mexico, Columbia, Peru and Ecuador[81].Semi-evergreen tree, loose and unruly, umbrella top with age and proper pruning, spreading and broad. sometimes ranging to 6m with greater spread. Masses of yellow flowers in axillary clusters followed by light brown pods[81].

      Full sunDrought-tolerant; suitable for xeriscaping. Grows well in areas that receive as little as 80−100 mm of annual rainfall. Supplemental irrigation will increase growth. Prefers full sun and well-drained soil in cultivated landscapes. Has a pH tolerance of 6.1−7.8[86].

      • Suited to large, open desert gardens where its architectural uniqueness can be featured.
      • Should not be planted in areas where its crown needs to be raised or pruned excessively[81].
      • Needs little to no supplemental water once established[81].


      ShrubsAcacia victoriae

      Bramble wattle
      Fabaceae
      AustraliaA multi-branched 3.5−4.5 m shrub that makes a good hedge if mass planted. It performs best in warm, dry conditions. Pale yellow flowers in two-to-three-inch racemes[76].Full sunPerforms best in warm, dry conditions[76]. It grows best in areas where annual daytime temperatures are within the range 12−32 °C, but can tolerate 5−38 °C[83]. Prefers a mean annual rainfall in the range 300−800mm, but tolerates 100−1,000 mm. Succeeds in most soils, being able to tolerate occasional short-term inundation. Tolerant of saline soils, yet prefers soils with a pH in the range 6−7.5, tolerating 5−8.5[83].The probable lifespan is 10−15 years[83].
      Shrubs/
      Grasses
      Atriplex spongiosa


      Pop saltbush
      Amaranthaceae
      AustraliaAnnual or short-lived perennial forb, grows up 40 cm tall or sprawling. Leaves are 1−4 cm long, 3–15 mm wide. Flowers are tiny (less than 3 mm across), male and female flowers on the same plant[84].
      Full sunStabilizing soil and/or make conditions more acceptable for the establishment by other species[51]. High salinity and drought tolerance.• High fodder quality and biomass production.
      • High water efficiency[51].
      Maireana sp.


      Band blue bushes
      Amaranthaceae
      AustraliaSucculent-like blue-white shrub[85].
      Full sun to partial shade
      Grows well in most soils, including alkaline clays. Prefers full sun but will grow in partial shade in desert areas[85]. Drought tolerant once established[85].• Makes an excellent container shrub and cut flower[85].
      Chrysopogon zizanioides


      Vetiver grass
      Poaceae
      IndiaStiff and erect stems, up to 2 m high, which can be trimmed to form neat hedges, stem flower heads with light purple flower, turning to mauve color later. It is sterile, so it is not invasive and has no weed potential[77].Full sun to partial Shade
      Tolerance to extreme climatic variation such as prolonged drought, flood, submergence and extreme temperatures (15–60 °C). It can tolerate precipitation levels as low as 300 mm and with up to 6-month drought[77].

      • Massive and penetrating root system that can grow very fast, up to 3–4 m in the first year.
      • Will continue to grow with deposited silt eventually forming terraces, if trapped sediment is not removed.
      • New shoots develop from the underground crown making vetiver resistant to fire, frosts, traffic and heavy grazing pressure[77].
      Eragrostis curvula


      Weeping lovegrass
      Poaceae
      Angola, Botswana, Burundi, Cameroon, Cape Provinces, Eritrea, Ethiopia, Free State, Kenya, KwaZulu-Natal, Lesotho, Malawi, Mozambique, Namibia, Northern Provinces, Swaziland, Tanzania, Zambia, Zaïre, ZimbabweDensely tufted, perennial grass. Its growth habit varies from erect to prostate, reaching a height of 30–120 cm. Stems can be slender or robust, with green to purple nodes, usually erect but sometimes bent at lower nodes. Leaves are dark green to blue-green. Basal leaf sheaths are keeled, strongly striate, usually hairless but sometimes silky hairy below and are typically straw-colored or purplish[88].Full sun
      Inhabits semi-arid subtropical grasslands and open scrublands, mainly on low-fertility, acidic sands and light-textured sandy loams, and particularly areas with significant elevation[88]. It is often associated with disturbed or badly managed areas. Grows in areas where annual summer rainfall is 400–1,000 mm and mean minimum and maximum temperatures are between 0 °C and 30 °C respectively[88].


      • Can become dominant when subject to heavy grazing pressure on disturbed land.
      • Growth rates tend to decline when competing with other plants.
      • Often suffers from fungal infections, mites and nematodes in areas where summer rainfall exceeds 700 mm and poor tolerance to flooding and standing water[89].


      Ground coverAmbrosia chamissonis


      Silver bur, ragweed
      Asteraceae
      Alaska, British Columbia, California, Chile Central, Chile South, Mexico Northwest, Oregon, WashingtonPerennial herb from long, creeping rhizomes; stems leafy, spreading-hairy, stout, branched just below the surface, forming large, lax clumps 20–160 cm tall. Leaves are succulent, mostly alternate, oblong to rounded in outline[90].Full sunPlants occur in moist to mesic coastal sandy or gravelly upper beaches and shorelines[90]
      • Attracts insects and birds.
      • Erosion control
      Artemisia californica


      California sagebrush

      Asteraceae
      California, Mexico Northwest
      Aromatic, subshrub usually 0.6–1.5 m tall but sometimes reaches 2.5 m, tend to be wide with many branches from the base. Upper branches are somewhat unbranched and the stems are whitish with appressed hairs, long, slender, leafy, and flexible[91].Full sunOccur on virtually all soil types except serpentine and on a variety of soil textures including depth loams, clayey loams, sandy loams, and loamy sands[90]. Drought tolerant, total annual precipitation ranges from about 25 – 60 cm, but can be higher. Requires full sun or nearly full sun[91]. California sagebrush can be compact if kept pruned and subjected to occasional pinching.• Provides habitat for many plant and animal species.
      • Leaves have been used by Native Americans for smoking, in sweat-houses, and various other purposes.
      • Said to stimulate the uterine mucosa, ensuring rapid childbirth and, if regularly consumed as a decoction prior to the onset of each menstruation, to prevent menstrual cramps and alleviate menopausal trauma.
      • Fresh or dried leaves were chewed to alleviate colds[91].
      Lasthenia glabrata


      yellowray goldfields
      Asteraceae
      California, Mexico Northwest
      An annual that grows up to 0.3 m, with daisy like yellow flowers. The species is a hermaphrodite[92].Full sunGrows in light sandy soils to heavy clay soils, vernal pools, low alkaline fields, hillsides etc, especially in grassland and alkaline marshes, up to 1,300 m. Prefers a sunny position[92].• Seeds are edible raw or cooked. They can be parched and eaten dry or made into a powder or cooked like a porridge[92].
      Mimulus aurantiacus


      Bush monkey flower
      Phrymaceae
      Southwestern North America, southwestern Oregon, California, Baja, Mexico
      Bushy evergreen shrub with narrow lanceolate dark green leaves and flowering stems with trumpet shaped flowers, grows up to 90–120 cm tall and 90 cm wide[93].Full sun to partial sunGrows in clay, loamy and sandy soils with a soil pH that ranges acidic to alkaline, in full to partial sun[93].• Pollinated by birds and bees.
      • Great choice for beds, borders and rock gardens and stone walls[93].
      Limonium californicum


      Western Marsh Rosemary
      Plumbaginaceae
      California, Mexico Northwest, Nevada, Oregon
      Half-hardy annual, Hardy perennial, half hardy perennial, 10–70 cm tall[94].Full sunFull sunlight. Light soils. Sandy soil. However, can survive in dry soils[94].• Yearly feed for perennials.
      • Cut perennials to the ground in autumn.
      • Propagate: divide perennials in spring or autumn[94].
      Alhagi graecorum
      Mannatree
      Fabaceae
      California, Mexico Northwest, Texas
      Perennial thorny shrub that grows up to 1m tall, woody at the base in older plants, glabrous or sparingly hairy, with long creeping solons; stems erect or ascending, much branched, twigs spiny at their tips. Leaves are alternate, simple, entire, elliptic to obovate, 7–20 mm long with short petioles 1–2 mm long and stipules 1 mm long. flowers are pink to maroon, approximately 10 mm long with a short slender pedicel[95, 96].
      Full sunOccurs on a range of soils from silty loams to clays as well as saline depressions[96]. It is adapted to alkaline to neutral soils.• Roots are used to treat kidney stones and diarrhea[95].
      Ground coverConvolvulus virgatus
      Morning glory
      Convolvulaceae
      Gulf States, Iran, Oman, Pakistan
      Under shrub forming a small bush up to 40 cm high and 60 cm wide; stems from a deep woody taproot, many, ascending, rigid, green, glabrous, sometimes spinescent at the tips, weakly divaricate. Leaves sessile lanceolate or linear-lanceolate, glabrous, acute or acuminate, corolla 1.3–2.1 cm, usually white, sometimes pinkish[97].Full sunFull sunlight. Good drainage. Soil pH 6–8. Regular watering. It may be necessary to trim leaves to display flowers[98].• Great container plants and indoor plants, and also perform well in rock gardens, sunny beds and borders, or gravel patios.
      • Best to prune in middle to late spring or once flowering has completed[98].
      Crotalaria aegyptiaca

      Rattlepod
      Fabaceae
      Egypt, Gulf States, India, Oman, Palestine, Saudi Arabia, Sinai, Somalia, Yemen
      Perennial Shrub, reaches up to 60 cm high, multi-branched, leafless at flowering[100].Full sunDominates sandstone stone soils, but not found in limestone soils[101].• Grazed by camels and gazelles, but poisonous to sheep and goat.
      • Old dry plants may revive after rainfall[100].
      Dipterygium glaucum

      Blooming shrub
      Cleomaceae
      Djibouti, Egypt, Eritrea, Gulf States, India, Iran, Oman, Pakistan, Saudi Arabia, Somalia, Sudan, Yemen
      Erect woody herb up to 80 cm high with numerous basal branches, slim olive-green leathery leaves. Flowers small with tubercles, yellow with pink tinge in few-flowered racemes; fruit/winged, pale green[108].Full sunHabitat sandy soils[35]. Mostly grows on deep sands, also on saline sands[108].
      • Grazed by livestock and often very stunted
      • Often flowering and fruiting in the first year
      • Used to treat respiratory disease[108].
      Heliotropium curassavicum
      Salt heliotrope
      Boraginaceae
      Americas; from Canada to Argentina
      Perennial herb, glaucous green borage. Leaves blue-green, obovate cuneate slightly fleshy. Flowers in scorpioid cymes usually a pair of cymes, white with yellow tinge[74].Full sunHabitat: Fields[74]. Sandy soils with a pH in the range of 6.6–8.5. It has a high tolerance to drought and anthropogenic disturbance. It can withstand areas with a rainfall of 200mm[106].
      • Dried and powdered plant is added to water and drunk to combat fatigue.
      • Topically to treat headache and used internally to treat gonorrhoea and to increase lactation.
      • Macerate of the plant is used as ink
      • Leaf sap is applied to burns[74].
      Rhazya stricta Decne.

      Harmal
      Apocynaceae
      Afghanistan, Gulf States, India, Iran, Iraq, Oman, Pakistan, Saudi Arabia, West Himalaya, YemenEvergreen dwarf shrub with thick foliage. It is a glabrous, upright perennial plant with many branches emerging from the base. Flowers are bisexual, white[102,103]. Yellowish-green shrub, up to 70 cm high.Full sunGrows in depressions with silty and sandy soils sometimes forming a pure stand and moderately salt-tolerant[102].
      • Used as a bitter tonic, to treat sore throat and fever.
      • Used to treat liver ailments[102].



      Rhynchosia minima

      Jumby-bean
      Fabaceae
      Afghanistan, Gulf States, India, Iran, Iraq, Oman, Pakistan, Saudi Arabia, West Himalaya, YemenPerennial herb, prostrate-trailing herb growing up to several meters long with trifoliolate leaves, the terminal leaflet rhomboid. Fruit a 2-seeded brown pod constricted between the seeds[119].Full sun Grows in woodland and grassland, roadsides, disturbed areas and depressions with sandy-clayey soil[119].

      • Plant is used as food for making sweet meats.
      • Root is laxative, vermifuge. It is used in the treatment of diarrhea, dysentery.
      • Leaf-sap and the root are used to treat hemorrhoids.
      • Leaves are abortifacient, inducing contractions of the uterus[119].
      Senna italica

      Italian senna
      Fabaceae
      Africa to the Indian Subcontinent.Leafy low shrub, up to 0.60 cm high with numerous branches ending in terminal racemes of yellow flowers distinct by the orange spots at the bases of the leaflets; fruits falcate tuberculated pods with persistent styles[119].Full sunIt is found in Runnels, wadis and depressions[119]. Will tolerate moistly and very poorly draining soils[107].

      • Leaves, pods and unmatured seeds are used as purgative, decoction and maceration to treat stomach complaints, fever, jaundice, venereal diseases and biliousness.
      • Used as abortifacient and against intestinal worms.
      • Leaves fresh or dried or pulverized used to dress skin problems, burns and ulcers. Flowers are made into tea and used as purgative and to induce labor.
      • Root maceration are used to cure colic and influenza.
      • Boiled roots are used to dress wounds.
      • Root infusion is used as eye drops for sore eyes and for the treatment of indigestion, liver complaints, gall bladder, nausea, vomiting and dysmenorrhea. Young seeds are eaten as snacks or as vegetable[107].
      Tribulus arabicus

      Zahr
      Zygophyllaceae
      Gulf States, Oman, Saudi Arabia, Yemen.
      Perennial herb up to 70 cm high with grey-green leaves and with large yellow flowers (1.5–2.0 cm)[104].Full sunStabilizes sand and sand plains[109].• No records of its use[109].
      Pulicaria glutinosa

      False fleabean
      Asteraceae
      NE. Tropical Africa, Arabian Peninsula, S. Pakistan.
      Subshrub 20–45 cm high, densely branched. Branches glabrous, glutinous. Leaves linear, 10–30, 1–2 mm, margins strongly revolute, apex apiculate, glabrous, sticky[110].Full sunSalt-tolerant plants grow in exceptionally salt-rich soil or waters of high-salinity conditions, including marshy and coastal lands. Adapted to survive in the prevailing dry climate and high-salt soil conditions[111].• Several Pulicaria species are used to treat inflammation, diabetes, and gastrointestinal disorders.
      • Plants from the genus are also used as tonic, food-preservative, in perfumes, and as part of salad[111].
    • There are 374 native plant species that belong to 55 families[9,40,112]. A significant amount of desert ecosystems are usually rich in annual flora[8]. In Kuwait, annuals make up 68% of the native flora, herbaceous perennials 22%, shrubs, undershrubs 9% and there is only one tree species (Vachellia gerrardii)[8,9,40,113]. There are five plant community types that have been identified in Kuwait; Haloxylon salicornicum (Moq.) Bunge ex Bioss, Rhanterium epapposum Oliv. and Cyperus conglomeratus Rottb which are used for livestock grazing, Zygophyllum qatarense Hadidi and Panicum turgidum Forssk[9,40,112,114]. Plant communities in arid extreme environments have the capacity to respond to environmental change and sustain functions that can impact species turnover and can shift interactions and net biodiversity[115,116]. Perennial and annual plants have an interrelationship between them that create negative and positive interactions between these life forms[62,115,116]. One of these interactions is that perennial plants benefit from early seedling emergence that is balanced by the risk of seedling mortality by adult longevity, in comparison to annuals that are more likely than perennials to postpone germination to following seasons[117]. As annual plants development, flowering and seed yield communities, success in desert ecosystems is dependent on a variety of environmental factors: rain, temperature, relative humidity, soil moisture availability, seed location, seed size and seeds depth in the soil, day length, light, soil water content, soil organic matter availability and nitrogen recycling[116,118]. Another interaction is that perennial plants provide two important contributions towards annual success; they act as organic-matter donors creating fertile islands from self-resource due to their root activity improving the quality of their understory soil as well as their intensive canopy cover acting as a physical structure in the ecosystem, creating a wind barrier and as soil-moisture moderators by providing shade[62,116,118]. Therefore, a selection of perennial plants should be utilized in garden design to facilitate the success of annual plants that provide a seasonal aesthetic and also colourful accents.

    • There are varying methodologies towards vegetation establishment in urban landscapes either through using hardened seeding or direct seeding. Direct seeding offers a more cost-effective and highly efficient planting method. Several studies have been conducted in Kuwait towards desert habitat rehabilitation through direct seeding utilizing pretreatment to break seed dormancy[43,57,59,62,68,71,118]. A study conducted by Suleiman et al.[40] in Kuwait on plant enrichment in desert ecosystems used a desert seed mix of Eragrostis curvula, Ambrosia chamissionis, Artemisia californica, Lasthenia glabrata, Mimulus aurantiacus and Limonium californicum[43]. Kuwait native species, hardwood cuttings of Lycium shawii treated with Hormex (1,000 mg/LL(IBA)) using the quick dip method and treated seeds of Ochradenus baccatus, Rhanterium epapposum and Nitraria retusa, with 500, 750 and 1,000 mg/LL GA3, respectively, and untreated seeds of Peganum harmala were tested[62]. A 100% survival rate was observed in Lycium shawii, Nitraria retusa and Ochradenus baccatus, in comparison to Peganum harmala and Rhanterium epapposum, which had a survival rate of 97.8% over approximately a one-month period[62]. The recommended plant spacing for native plants (Table 3) is only a guideline based on previous research[43,62]. However, plant spacing is dependent on the space, aesthetic or whether it is grown in a square grid, rectangular grid, or triangular spacing.

    • Combating natural habitat loss and enhancing plant diversity by improving urban landscapes in Kuwait is a critical issue that needs to be addressed to ensure the survival of native flora and fauna. Conservation and rehabilitation efforts should not be focused solely on desert areas and nature reserves as they fail to address the issues of habitat fragmentation. As the main constraints faced by plants are Kuwait's harsh desert environmental conditions, poor soil and water scarcity, there needs to be a shift away from the current plant palette that utilizes plant species that have become invasive and have high water and fertilizer requirements. The use of native and xeric plant species that have been recommended in this paper will assist in providing guidance in the future when creating more sustainable urban green landscapes that offer a variety of ecosystem services.

    • A special thanks to Proceso T. Ramos Jr. for contributing to the identification and confirmation of native plant species.

      • The author declares that there is no conflict of interest.

      • Copyright: © 2022 by the author(s). Published by Maximum Academic Press, Fayetteville, GA. This article is an open access article distributed under Creative Commons Attribution License (CC BY 4.0), visit https://creativecommons.org/licenses/by/4.0/.
    Figure (3)  Table (4) References (122)
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    Almutawa AA. 2022. Native and xeric plant recommendations for urban landscapes in Kuwait. Technology in Horticulture 2:7 doi: 10.48130/TIH-2022-0007
    Almutawa AA. 2022. Native and xeric plant recommendations for urban landscapes in Kuwait. Technology in Horticulture 2:7 doi: 10.48130/TIH-2022-0007

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